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Project Name Data (Last Modified On 7/23/2013)

Flora Data (Last Modified On 7/23/2013)
Species Macleania insignis Mart. & Gal.
PlaceOfPublication Bull. Acad. Roy. Sci. Bruxelles 9: 531. 1842.
Note TYPE: Mexico: Veracruz, Galeotti 1827 (BR, holotype, not seen; W, isotype, not seen; photo, GH, US).
Synonym Thibaudia laurifolia Mart. & Gal., Bull. Acad. Roy. Sci. Bruxelles 9: 530. 1842. TYPE: Mex- ico; Oaxaca, Galleotti 1814 (BR, holotype not seen; W, isotype not seen; photo, GH, US). Macleania ovata Klotzsch, Linnaea 24: 20. 1851. TYPE: "in America centrali prope Ver- aguam," Warszewicz (B, holotype, not seen; photo, F). Macleania cordata var. linearifolia Donnell Smith, Bot. Gaz. (Crawfordsville) 16: 12. 1891. TYPE: Guatemala: Pasamala, Tuerckheim 1332 (US, holotype; photo, NY). Macleania linearifolia (Donnell Smith) A. C. Smith, Contr. U.S. Natl. Herb. 28: 368. 1932. Macleania compacta A. C. Smith, Contr. U.S. Natl. Herb. 28: 369. 1932. TYPE: Mexico; Veracruz; near Jalapa, Pringle 7870 (GH, holotype). Macleania insignis var. linearifolia (Donnell Smith) Standley & L. Wms., Fieldiana, Bot. 31: 172. 1965.
Description Shrubs often epiphytic 0.5-2 (-4) m tall; branchlets terete, light to dark brown; bark peeling, puberulous; young twigs glabrous to puberulous. Leaf blades ovate to lance ovate or rarely lance oblong to oblong, (2-)3-8(-12) cm long, (1-) 1.5-4 cm wide, basally subcordate, rarely rounded, apically obtuse to acute, entire and slightly revolute, glabrous on both surfaces; 3-5(-7)-pli- nerved with the secondary nerves originating near the base, ascending, the mid- vein slightly impressed above and elevated beneath; petioles subterete, rugose, glabrous or rarely puberulous, (1-)2-4 mm long. Inflorescence axillary, subfas- ciculate, (1-)2-3-flowered, basally circumscribed by a few, minute, ovate, ciliate bracts 1.5-2 mm long; pedicels rugose, glabrous or rarely puberulous, (4-)8-16 mm long, bracteoles basal, minutely ciliate, triangular, 1-2 mm long. Flowers with the hypanthium glabrous or minutely to moderately appressed short pu- bescent, prismatic, 3-S5 mm long, 3-4 mm in diameter, broadly winged from the base to the sinuses, the wings 0.3-0.5 mm wide; calyx limb ascendent to erect, 1.0-1.5 mm long, the lobes 5, apiculate, narrowly triangular to deltoid, (0.4-) 0.8-2.0 mm long, the sinuses either flat or acute, more or less contiguous; corolla elongate urceolate or prismatic, (14-)20-25 mm long, 4-5 mm in diameter, con- tracted at the throat, externally glabrous to slightly puberulous especially dis- tally, distally sparsely to densely villous internally, the lobes acute, spreading, 1.5-3.0 mm long; stamens 10, 8-12 mm long, the filaments dark brown, glabrous, connate 3-5(-7) mm long, the anther sacs 4.0-5.5 mm long, the single, cylindric- conical tubule 2-3 mm long opening by a broad introrse cleft more than 1/2 as long as the tubule; style exerted, the stigma peltate.
Habit Shrubs
Distribution As interpreted in this account, Macleania insignis ranges from the moun- tains of southern Mexico southward into Panama.
Note The present treatment is a drastic reduction from the number of species recognized in Smith's (1932) monograph of the genus. The representatives of this genus with connate filaments in southern Central America have been re- ferred to M. ovata Klotzsch. Macleania ovata has been generally differentiated from the more northern M. insignis by its longer and broader, deltoid calycine lobes which form acute basal sinuses with the adjacent lobes. Macleania in- signis supposedly had minute calycine lobes less than 1.5 mm long forming a broad, flat sinus between adjacent lobes. There is much greater variability in calycine lobes than was apparent in the relatively few specimens available to Smith, and there is an apparent continuum between the extremes. Hence, there is no way to recognize two taxa based on calycine lobes. Yeo (1967) suggested that M. ovata might be best regarded as a geographical subspecies of M. insignis but he did not make the combination. The calycine variation does not suggest geographical subspecies since apiculate-lobed plants are found at least as far south as Costa Rica and deltoid, larger lobed plants are to be found in southern Mexico along with the apiculate-lobed plants.
Specimen PANAMA: Moist upper regions of Cerro Campana, 1000 m, Allen 2406 (A).
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