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(Last Modified On 7/22/2013)
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(Last Modified On 7/22/2013)
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Species
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Cavendishia complectens Hemsl.
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PlaceOfPublication
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Biol. Centr. Amer., Bot. 2: 272. 1881.
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Note
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TYPE: Costa Rica, Endres 257 (K, holotype; photo of holotype, DUKE, NY; W, isotype).
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Description
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Epiphytic and terrestrial shrubs, 1-1.5 m tall, glabrous throughout; stem base to about 3 cm in diameter; mature branch erect or pendent, terete to bluntly angled; bark grayish to reddish brown. Leaves usually subsessile, amplexicaul, drying olive brown or dark green, oval to semiorbicular, (5-)8-16(-18) cm long, (4-)5.5-10(-13) cm wide, basally cordate with the basal lobes rounded, usually imbricate, apically obtuse to broadly rounded, rarely bluntly cuspidate; pin- nately veined, the midrib impressed above, raised and conspicuous beneath, lateral nerves usually raised on both the surfaces, anastomosing to form a weak, collective marginal nerve, the veinlets slightly raised and obscure to conspicu- ous; petioles subterete, rugose, to 5 mm long. Inflorescence 1(-3) rachises per axil, the largest rachis rarely once branched at the base, 10-27(-60)-flowered, the rachis terete to bluntly angled, swollen at each floral node, 2-9(-20) cm long and 1-2 mm in diameter at anthesis, increasing to 3 mm diameter post- anthesis, pinkish green at anthesis, later turning purplish; floral bracts smooth or becoming ribbed after drying, elliptic to ovate lanceolate, rarely subor- bicular, basally narrowed and somewhat auriculate, apically acute, (10-)11- 16 (-20) mm long, (5-) 7-10 (-17) mm wide, dark pink to red, marginally scarious and glandular fimbriate, ascending at anthesis but later spreading or perpendicular to the rachis; pedicels minutely striate or rugose, (2-)4-9(-11) mm long, 0.5-1 mm in diameter, pinkish green at anthesis but later turning purplish; bracteoles oblong to lanceolate, (1-) 3-4 (-7) mm long, (0.5-) 1-2.5 (-4) mm wide, basally slightly auriculate, apically rounded, marginally glandular fimbriate, dark pink. Flowers with the calyx (4.5-)6-8(-10) mm long, the tube hemispheric, basally not enlarged, smooth or minutely muricate, (1.5-)2-3.5(-6.5) mm long, 2.5-6 mm in diameter, purplish green, the limb campanulate, coriaceous, (2.5-)4-5(-5.5) mm long including the lobes, pale green, the lobes reniform, broadly ovate, rarely oblong, imbricate, (1.5-)2-3(-4) mm long, (1.5-)3(-4.5) mm broad, glandular fimbriate, whitish green, flaring at anthesis, later conni- vent and tightly curling around the base of the style; corolla cylindric or bottle- shaped, narrowed at the throat, (7-)8-10(-12) mm long, 4-6.5 mm in diameter at the widest part, white at the base, purplish black in the basal 1/4, the mid- section green, the limb white, the lobes oblong, obtuse, 2-2.5 mm long, white; stamens 7-8.5 mm long, the filaments white, alternately 2-3 mm and 3.5-4 mm long, with a few scattered distal hairs, the anthers orange including tubules al- ternately 4.5-6.5 mm and 5.5-7 mm long, the thecae 2-4 mm long; style 7-8.5 mm long, distally flaring. Berry 9 mm in diameter.
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Habit
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shrubs
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Note
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Cavendishia complectens is one of the easiest species to recognize because of its cordate, amplexicaul leaves. It has been collected a few times in eastern Darien Province, Panama, and in Choco, Colombia, but is to be expected from western Panama also. It is common in the mountains of the Central Valley of Costa Rica and occurs rarely in Nicaragua.
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Specimen
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DARIEN: Cerro Pirre, Bristan 596; Duke 5321, 6103 (all MO). Serrania de Pirre, head- waters of Rio Escucho Ruido, ca. 16 km N of Alto de Nique, 1530-1550 m, Croat 37936 (NY). W slope of Cerro Pirre, 1000 m, Duke 6595 (MO). Cerro Campamiento, Duke 15728(1) (MO). Cerro Pirre, 770-1380 m, Duke & Elias 12751 (F, MO, NY). Cuasi-Canai Trail on Cerro Campamiento, Kirkbride & Duke 1224 (MO, NY). Causi-Cana' Trail between Cerro Campamiento and La Escalera, Kirkbride & Duke 1339 (MO, NY). N slopes of Cerro Pirre, 700-950 m, Mori & Kallunki 5465 (NY). Crest of Cuasi-Cana' Trail, 1690 m, Terry & Terry 1577 (A, F, MO).
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