(Last Modified On 7/22/2013)
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(Last Modified On 7/22/2013)
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Species
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Cavendishia pubescens (H.B.K.) Hemsl.
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PlaceOfPublication
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Biol. Centr. Amer., Bot. 2: 273. 1881.
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Synonym
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Thibaudia pubescens H.B.K., Nov. Gen. Sp. PI. 3: 273. 1818. TYPE: Venezuela: near Caracas, Humboldt & Bonpland (B, lectotype; not seen, presumably destroyed; P, isolectotype not seen, photo of isolectotype, DUKE, F, NY). Proclesia pubescens (H.B.K.) Klotzsch, Linnaea 24: 33. 1851. Thibaudia septemnervia Willd. ex Walp., Ann. Bot. Syst. 2: 1087. 1852, as syn., nom. illeg. Art. 34. Thibaudia cinnamomifolia Willd. ex Walp., Ann. Bot. Syst. 2: 1087. 1852, as syn., norn. illeg. Art. 34. Chupalon pubescens (H.B.K.) Kuntze, Rev. Gen. PI. 2: 383. 1891.
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Description
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Epiphytic and terrestrial shrubs, 1-3 m tall; mature branches subterete or bluntly angled, ridged, moderately short pilose, reddish brown when dry; im- mature branches and twigs of the new growth flattened and coarsely angled, ridged, densely short pilose with whitish or grayish trichomes ca. 0.5 mm long. Leaves elliptic to lanceolate, (5.5-) 14-20 cm long, (1.5-)3.5-9 cm broad, basally cuneate, obtuse or rounded, apically long acuminate with a blunt tip or rarely acute, grayish green when dry, densely puberulent on both the surfaces with soft, white trichomes, persistent beneath, often glabrate above; 5 (-7)- plinerved, the midrib impressed above and prominently raised beneath, the lateral nerves impressed above but plane distally, raised and prominent be- neath, the veinlets plane to slightly raised but obscure on both sides; petioles subterete, rugose, coarsely ridged, (6-)9-15 mm long, 2-3.5 mm in diameter, densely short pilose. Inflorescence 7-11-flowered, encircled at the base by densely short-pilose bracts; rachis bluntly angled, striate, rugose, short pilose with persistent, densely white hairs, sometimes fimbriate, 1-2.5(-4.5) cm long, 2-3 mm in diameter, pink, rarely with basal cartilaginous teeth; floral bracts muricate, short pubescent both abaxially and marginally with densely white hairs, elliptic to oblanceolate, basally narrowed, apically rounded, 17-30 mm long, 7-13(-17) mm wide, pale pinkish green or whitish green, usually with small, red, clavate glands abaxially; pedicels bluntly angled, densely short pilose, often fimbriate, 6-14 mm long, 1-1.5 mm in diameter, pink but turning green; bracteoles linear to aristate, rarely oblong, 2-4.5 mm long, to 1 mm wide, densely pubescent and glandular fimbriate. Flowers with the calyx pubescent with densely matted white or tan trichomes, glandular fimbriate, 5.5-6.5(-9) mm long, the tube cylindric, 2.5-4 mm long, not basally enlarged, green, the limb cam- panulate or spreading, (2.5-)3.5 mm long, whitish to pinkish green, the lobes ob- long, broadly triangular, obtuse or broadly rounded and apiculate, 1.5-2(-3) mm long, connivent after anthesis, pinkish green, the sinuses obtuse; corolla cylindric or bottle-shaped, slightly constricted basally, narrowed to the throat, (14-)17-19 (-22) mm long, 8 mm in diameter, white or in age tinted with pink, densely short pilose with white or tawny trichomes, the lobes triangular, wide spreading at anthesis; stamens 11-15.5 mm long, the filaments alternately 2.5-4 mm and 3.5-6 mm long, white, the anthers including the tubules alternately 8-11 mm and 9.5-13 mm long, brownish orange, the thecae 2.5-4 mm long; style glabrous or rarely distally pilose, 13.5-17 mm long. Berry densely puberulent, 8-14 mm in diameter.
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Habit
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shrubs
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Distribution
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Cavendishia pubescens is widespread in the Andes of South America from Bolivia to Venezuela. Its most northwestern location is Volcan Chiriqui, Panama, where it is common on the western side of Volcaln Chiriqui along the steep roadside slopes northwest of El Hato.
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Note
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It is readily distinguished by pale pink- ish to whitish green floral bracts, densely short pilose white corollas, densely pilose to almost wooly calyces, and by persistently puberulent leaves and fruits. The densely matted hairs of the calyx are in themselves enough to distinguish the taxon. Cavendishia pubescens is closely related to C. crassifolia. Several collections from the area between El Hato and Cerro Punta in the vicinity of Volcan Chiriqui suggest the possibility of hybridization between the two species. Such collections are noted below with an asterisk.
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Specimen
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CHIRIQUI: Road to Cerro Pando, 6 km NE of El Hato, 1840 m, Correa 1384 (DUKE). Road to Bambito, beyond the lianos, 1840 m, Correa & Lazor 1414 (DUKE, MO). Boquete, Davidson 649 (A, F, MO, US). 1.5 mi S of Cerro Punta, 1850 m, Graham 288 (GH, MICH). Road NW of El Hato towards Costa Rica, 1690 m, Luteyn 853 (DUKE, MO). Pasa Ancho, 1840 m, Luteyn 3053* (DUKE), 3054 (DUKE, F, MO, NY). Road NW of El Hato towards Costa Rica, 1690 m, Luteyn 3055 (CAS, DUKE, F, GH, LL, MICH, MO, MSU, NY, UC, US, WIS). Road 2 km above Bambito towards Cerro Punta, Luteyn & Wilbur 4652* (DUKE, F, GH, MICH, MO, NY, US). Boquete, Seemann 1613 (BM, K). Just S of Boquete, Stern et al. 1959 (MICH, MO, US). At opening to canyon to Bambito, 1540 m, Tyson 5862* (FSU, MO). NE of Cerro Pando, 1500 m, Wilbur et al. 11015 (CAS, DUKE, F, FSU, G, GH, LL, MICH, MO, MSU, NY, PMA, UC, US, WIS). Slopes between Nueva Suiza and Bambito, 1690 m, Wilbur et al. 11878, 11879* (both DUKE).
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