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(Last Modified On 7/23/2013)
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(Last Modified On 7/23/2013)
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Species
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Disterigma humboldtii (Klotzsch) Niedenzu
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PlaceOfPublication
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Bot. Jahrb. Syst. 11: 224. 1889.
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Synonym
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Vaccinium humboldtii Klotzsch, Linnaea 24: 57. 1851. TYPE: S. America, Humboldt 2065 (B, holotype, not seen and presumably destroyed). Vaccinium pachyphyllum Hemsl., Biol. Centr. Amer. Bot., 2: 275. 1881. TYPE: Costa Rica, Endres 154 (K, holotype, not seen; photo of holotype, NY). Disterigma pachyphyllum (Hemsl.) Blake, J. Wash. Acad. Sci. 16: 365. 1926. Disterigma mayanum Lundell, Wrightia 5: 85. 1975. TYPE: Guatemala; Dept. Baja Verapaz, Lundell & Contreras 18936 (LL, not seen).
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Description
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Bushily compact to straggly, terrestrial or epiphytic shrubs (2-)5-10(-12) dm tall; branchlets ridged and grooved and indistinctly 3-angled to irregularly 4-angled, typically setose with tawny to chocolate brown trichomes mostly 0.5 to 1.2 mm long or moderately to densely puberulous, becoming glabrate and terete; internodes short on lateral branchlets, usually less than half as long as the nearest leaf, often only 1-2 mm long, but 1-2 times as long as the nearest leaves on fast growing leading shoots. Leaves coriaceous, entire, ovate to ellip- tic to almost orbicular, 6-10(-16) mm long, 4-6(-9) mm wide, apically obtuse to rounded, basally rounded, glabrate or with a sparse scattering of appressed, clavate trichomes 0.2 mm long or less on the lower surface, commonly with a dense but short, apical tuft or marginal cilia, the midvein slightly impressed above, the venation otherwise completely obscure both above and beneath, dark green above and pale green beneath, this distinction persisting when dried; petioles 1 (-2) mm long, sparingly to moderately spreading short pubescent. Inflorescence of solitary flowers to 2-3 per axil; pedicels ca. 0.5-1(-1.5) mm long; bracts 2-3, basal, broadly clasping, 2-3 times broader than long; bracteoles glabrous, 2, broadly clasping, oblong, truncate, 3-4 mm long and nearly as broad, borne just below and embracing the base of the flower. Flowers with the hypanthium 4-angled, campanulate, glabrous, ca. 1 mm long, 1-1.5 mm in diam- eter; calyx lobes 4, erect to slightly spreading, lance deltoid, acute, 1.5-2.5 mm long, entire and eciliate, 1.5-2.5 mm long, glabrous or sparsely appressed pubescent; corolla white or white and deeply suffused with rose or pink, the tube cylindric, 6-8 mm long, 2-4 mm in diameter, glabrous internally and glabrous to sparingly appressed pilose externally, the lobes 4, erect to horizontally spread- ing, ca. 1 mm long, deltoid, acute; stamens 8, about as long as the corolla, the filaments 3.5-5 mm long, flattened moderately to densely hyaline setose, the anthers slightly granular, ca. 1.5-2 mm long, each theca tapering into a slender tubule 2.5-3.5 mm long, style glabrous, slightly exserted beyond the corolla lobes.
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Habit
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shrubs
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Note
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As is suggested by the description, there is much variability in this species and yet none of it merits formal recognition.
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Distribution
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known from Guatemala, Costa Rica, Panama (Bocas del Toro, Chiriqui and Veraguas) and northern South America.
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Specimen
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BOCAS DEL TORO: Robalo Trail, northern slopes of Cerro Horqueta, 6000-7000 ft, Allen 4986 (F, MO, US). CHIRIQUI: Bajo Mono-Robalo Trail, W of Cerro Horqueta, 5000-7000 ft, Allen 4845 (F, MO, NY, US). Bajo Chorro, Boquete District, 6000 ft, Davidson 92 (F, MO). Upland pastures, Las Nubes, 7200 ft, Luteyn 3039 (DUKE). Valley of the upper Rio Chiriqui Viejo, White 25 (MO). Bajo Chorro, 1900 m, Woodson & Schery 633 (MO, NY), 684 (MO, NY). VERAGUAS: Cerro Tute, ca. 10 km NW of Santa Fe, on ridgetop in cloud forest above 1000 m, Mori et al. 7597 (MO).
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