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Project Name Data (Last Modified On 7/24/2013)

Flora Data (Last Modified On 7/24/2013)
Species Themistoclesia costaricensis Luteyn & Wilbur
PlaceOfPublication Brittonia 29: 265. 1977.
Note TYPE: Costa Rica, Luteyn et al. 4407 (DUKE, holotype; F, MO, NY, isotypes).
Description Pendent, epiphytic shrub with branches to 0.5 m long; stems terete, minutely striate, tan to reddish brown when dry, hispidulous with reddish brown trichomes 0.6-0.8 mm long, glabrate with age. Leaves distichous, flat or concave, ovate, (1.6-)2-2.5 cm long, (0.8-)1-2 cm wide, basally rounded or subcordate, apically acute or short acuminate, entire, hispidulous above especially near the margins, glabrous beneath, brown above and olive green or light brown beneath when dry; venation obscurely 5-plinerved, the midrib slightly impressed proximally, raised distally above, slightly impressed but inconspicuous beneath, the lateral nerves and the veinlets slightly raised but inconspicuous above, totally obscure beneath; petiole subterete, densely puberulous to pilosulose, 1-1.5 mm long. Inflorescence of axillary, solitary flowers with 4 minute basal inflorescence bracts; floral bract ovate, ciliate, ca. 1.5 mm long; pedicel terete, glabrous, 1-2 mm long, completely concealed by the bracteoles; bracteoles 2, ovate to hemi- spheric, distally hispidulous. Flowers with the hypanthium glaucous, copiously pilosulose with translucent or reddish tinged trichomes 1-1.2 mm long, the tube obconic, bluntly 5-angled, ca. 3 mm long; calyx limb 1-1.5 mm long including apiculate lobes; corolla cylindric, narrowed at the throat, 5-6 mm long, ca. 2.2 mm in diameter near the base, pilosulose throughout, white, the lobes ca. 1 mm long; stamens 10, ca. 5 mm long, the filaments alternately 2.8-3 mm and 3-3.2 mm long, pilosulose over the entire length, the anthers ca. 2.5 mm long, the tubules slightly longer than the thecae, the theca with a minute, 0.1 mm long, fingerlike appendage protruding from the base; style 5-6 mm long, glabrous. Berry not seen.
Habit shrub
Distribution known from Serrania del Darien and from Costa Rica.
Note The identity of the Panamanian specimen is not certain as the ma- terial is immature, and there are discrepancies between it and specimens from the type locality along the Rio Grande de Orosi in central Costa Rica. These differences include cauline trichomes 0.8-1.2 mm long vs. 0.6-0.8 mm long, leaves to 3 cm long vs. 1.6-2.5 cm long, bracts and bracteoles glandular fimbriate vs. ciliate, and pedicels sparsely pilosulose vs. glabrous.
Specimen DARIEN: summit of Pico Tacarcuna, highest point of the Serrania del Darien, 1850 m, Gentry et al. 16911 (NY).
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