|
(Last Modified On 7/24/2013)
|
|
|
|
(Last Modified On 7/24/2013)
|
|
Species
|
Themistoclesia pentandra Sleumer
|
|
PlaceOfPublication
|
Bot. Jahrb. Syst. 71: 392. 1941.
|
|
Note
|
TYPE: Costa Rica, Prov. de Cartago, La Estrella, Standlley 39374 (US, holotype).
|
|
Description
|
Epiphytic, often slender and straggly shrub 3-6(-15) dm tall with ascendent branches; stems terete, striate, fissured, grayish to brown or reddish brown, moderately to densely spreading short pubescent or pilosulose with hyaline trichomes 0.2-0.6 mm long, glabrate with age. Leaves spirally arranged, elliptic to oblong elliptic, (1.5-)2.0-3.5(-4.5) cm long, (0.8-)1.2-2.0(-2.5) cm wide, basally cuneate, less commonly rounded, apically acute to obtuse, entire, moderately but inconspicuously spreading pilosulose to puberulous on both the surfaces when young, becoming glabrate or more sparsely pubescent, the lower surface also inconspicuously moderately glandular strigillose, obscurely 3-5- plinerved, the midvein and the principal veins slightly elevated above and be- neath, the lateral veins obscurely reticulate on both the surfaces; petiole flat- tened above, pilosulose to puberulous, rugose, 1-2(-3) mm long. Inflorescence of axillary, solitary or very rarely paired flowers; bracts 2 or 4, appressed, trian- gular 1-1.5 mm long; bracteoles 2, appressed, lanceolate, puberulous 1.2-1.5 mm long; pedicels terete, puberulous 1.5-2.5 mm long. Flowers with the hy- panthium 3.5-4 mm long, conspicuously 4-5-winged, the wings conspicuously ciliate with hyaline trichomes 0.2-0.4 mm long, these trichomes occurring else- where on the surface, the surface of the hypanthium sparingly to moderately beset with minute, glandular trichomes; calyx limb 0.8-1.0 mm long including the minute apiculate lobes; corolla sharply pentangular, tapering sharply to the throat, 3-4.5 mm long, basally 2.0-2.5 mm in diameter, moderately hyaline pilosulose throughout, both internally and externally, light orange with a pale yellow tip, the lobes 4-5, narrowly triangular, acute, 1.0-1.5 mm long, erect or the tips slightly spreading; stamens 5-10, 3.2-3.5 mm long, equal or nearly so, about as long as the corolla tube; filaments glabrous or sparingly ciliate, ca. 1.5 mm long, flattened basally, the anther sacs attached medially, the thecae slightly granular, 1.0-1.2 mm long, the tubules smooth, conical, 1.5-1.8 mm long; stigma truncate, the style 3-3.5 mm long or about as long as the corolla tube. Fruit 4-5-winged, the wings 1-2 mm wide, pale greenish, 5-7 mm long.
|
|
Habit
|
shrub
|
|
Note
|
Themistoclesia pentandra has been rarely collected and specimens seen have been incomplete. It is possible that more than one species is included under this binomial. Most recent collections have 8 or 10 stamens while the original description and indeed the specific epithet indicates 5. The inclusion of Duke & Elias 13688 from Darien is especially suspect due to the following differences from all other specimens of T. pentandra; the calyx is glabrous rather than ciliate, the corolla is glabrous externally and minutely papillose or short puberulous within in contrast to being externally and internally pilosulose, and its stamens, including the filaments, thecae and tubules are all shorter. These differences are significant but we are reluctant to name another species based upon a single, rather incomplete specimen.
|
|
Specimen
|
CHIRIQUI: N of San Felix on Cerro Colorado Copper Mine road along continental divide, 5000-5500 ft, Mori & Kallunki 5984 (MO). COCLE: Cloud forest on slopes of Cerro Pil6n near El Valle, 700-900 m, Duke 12080, 12085 (both MO). Tropical wet forests, La Mesa, 4 km N of El Valle, 875 m, Nee & Hale 9612 (MO). DARIEN: Cloud forest and/or mossy forest, Cerro Pirre, 2500-4500 ft, Duke & Elias 13688 (DUKE, MO).
|
|
Tag
|
|
|
|
Project Name
|
|
Tag
|
|
|
|