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Project Name Data (Last Modified On 9/30/2013)

Flora Data (Last Modified On 9/30/2013)
Species Mucuna holtonii (Kuntze) Moldenke
PlaceOfPublication Phytologia 1: 7. 1933.
Synonym Stizolobium holtonii Kuntze, Rev. Gen. P1. 1: 207. 1891. SYNTYPES: Costa Rica, Kuntze (?NY, not seen) and Colombia, Holton 971 (K, not seen). Mucuna andreana Micheli, Morot. J. Bot. 6: 146, tab. 5, 6. 1892. TYPE: Colombia, Andre 1978, not seen. Also cites Holton 971, not seen.
Description High climbing vine, stems slender, slightly angled, minutely tomentulose, tar- dily glabrescent. Leaves trifoliolate, the leaflets about equal, 9-15 cm long, 5-9 cm wide, apically short acuminate, mucronate, manifestly costate, the terminal leaflet with 5-6 major veins on each side, the lateral leaflets strongly oblique, chartaceous, velutinous, later glabrescent especially above, with only a few hair bases remaining; petiolules 7-10 mm long, tomentose, glabrescent and turning black; petioles mostly shorter than the leaves, tomentose, glabrescent; stipules caducous; stipels ca. 3 mm long. Inflorescences pendant, contracted, racemes with peduncle elongate, to 5 m long, slender, flexuous, minutely tomentose ve- lutinous, the raceme enclosed in bud by gray velutinous, papery deciduous bracts and forming a globose 'bud' ca. 5 cm across, sinuous; pedicels velutinous, 15-18 mm long upon inflorescence opening, becoming 5 cm long at anthesis and stout in fruit. Flowers greenish or yellowish, the calyx deciduous, cupular, velutinous outside with appressed, ascending gray or golden hairs, ca. 10-15 mm long, pilose inside, the teeth small, deltoid umbonate; corolla ca. 4 cm long, drying dark, the standard 18-22 mm long, the wings and keel narrowly tubular, exserted about 1/2 their length from the standard; anthers basally barbate with elongate hairs, the stigma capitate, a tuft of ascending hairs. Fruit oblong, flat, 1-6 seeded, 14-25 cm long, 5 cm wide, the margin conspicuously undulate between the seeds, softly villous outside, lustrous within and falsely septate between the seeds; seeds dis- coid, black, 2-3 cm wide, ca. 0.8 cm thick, the hilum linear around 3/4-4/5 of the periphery, lighter colored.
Habit vine
Note Mucuna holtonii has leaves velutinous on both sides until time of flowering. The inflorescence is greatly contracted, appearing in bud as a ball at the end of a long, slender, pendant peduncle, and later with the flowers on elongate and slightly bent pedicels. The legumes are flat with undulate margins. The legume lacks stinging hairs, but there may be some on the calyx. This species is found in Panama from sea level to at least 1,500 m, occurring in forests and in areas of disturbance. It ranges from Mexico (Chiapas), to Co- lombia. It is apparently closely related to Peruvian and Ecuadorean material going under the name M. elliptica Ruiz & Pav6n, which differs in having seeds slightly thicker and more convex, and in the legumes having stinging hairs. The appear- ance of the legume and also of its indumentum is much like that of M. elliptica.
Elevation sea level to at least 1,500 m
Specimen BOCAS DEL TORO: Banks of Changuinola River, Dunlap 441 (F, US). Changuinola to 5 mi S at junction Rios Changuinola and Terebe, 100-200 ft, Lewis et al. 943 (GH, MO, US). Almirante Bay, Wedel 8 (MO, US). CANAL ZONE: Road C-16 NW of Pedro Miguel, Croat 12255 (MO). Pipeline road 3 mi N of Gamboa gate, Croat 13951 (MO, SCZ). Madden Dam, Dwyer 1965 (MO, PMA, US). Between Chilibre and Madden Dam on Transisthmian Highway, Dwyer 8396 (GH, MO). Chagres River 3 mi above Gamboa Bridge, Kennedy et al. 2304 (F, MO). Pipeline Road 2 km NW of Gamboa, 50 m, Nee 7853 (MO, US). Gatuncillo, Piper 5136 (US). Gamboa, Pittier 4788 (US). COCLE: Lim6n, N of Alto Calvario, Folsom 5848 (MO). COL6N: Quebrada Bonita, 13 km NW of Buena Vista, 3 km NW of Salamanca, 100-170 m, Nee 7632, 9075 (both MO). DARIEN: Ascent of Cerro Pirre from Rio Pirre S of El Real, 100-600 m, Duke 5286 (MO). Between Rio Perrecenega and Rancho Frio, 500- 2500 ft, Duke & Elias 13877 (MO). Rio Areti, Duke & Nickerson 14925 (MO). Rio Pirre near town of Pirre, Gentry & Clewell 6922 (MO). Rio Pucro below Pucro, Gentry & Mori 13531 (MO). Quebrada Camachimuricate cerca casa de Bartolo, Kennedy 2840 (F, GH, MO). Cand, 1750 ft, Stern et al. 482 (GH, MO). VERAGUAS: Above Santa Fe on slopes of Cerro Tute, 1200-1400 m, Gentry 6280 (MO).
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