8. Quercus pagoda Raf. (cherrybark oak)
Quercus falcata Michx. var. pagodifolia Elliott
Pl. 414 i; Map
1846
Plants trees to
40 m tall. Bark medium to dark gray, divided into square blocks or plates, the
inner bark orange. Twigs 2–4 mm wide, dark brown, densely pubescent with
branched, spreading hairs when young, sometimes becoming more or less glabrous
with age. Buds 5–7 mm long, brown, pubescent or the lower scales glabrous
except along the margins. Petioles 27–45 mm long. Leaf blade 12–19 cm long,
10–15 cm wide, broadly obtuse or rounded-obtuse to nearly truncate at the base,
divided 70–80% of the width, the lobes (2)3 or 4 per side, these evenly spaced,
the median lobes usually the largest, the basalmost lobes 1.0–2.5 cm above the
blade base in most or all leaves; well-developed lobes 15–30 mm wide, narrowly
triangular, rarely almost oblong, usually tapered apically, occasionally
rounded-obtuse or acute, undivided or with a strong tooth (rarely a secondary
lobe) on the lower margin, each with 1–4 bristles 3–4 mm long (the whole blade
with 10–20 marginal bristles), the strongest secondary veins reaching the
margin at the tips of the lobes and ending in bristles, others reaching toward
sinuses and turning aside before reaching the margin; the upper surface dull or
somewhat shiny, with small branched spreading hairs when young, by midsummer
usually with only scattered hairs near the midvein, the undersurface green or
white, with moderately to densely crowded 7–11-rayed, spreading hairs and often
also inconspicuous, unbranched, appressed hairs over the whole surface, usually
felty to the touch, the vein axils with tufts of 5–15-rayed, often stalked
hairs. Acorn cups 6–8 mm long, 17–18 mm wide, covering 30–60% of the nut,
bowl-shaped, the inner surface smooth, densely hairy, the outer surface with
the scales thin and plane or weakly convex-thickened, pubescent. Nuts 10–12 mm
long, 11–12 mm wide, depressed-ovoid, without distinct concentric grooves
around the tip. April–May.
Scattered in the
Mississippi Lowlands Division and adjacent portions of the Ozarks and Ozark
Border (southeastern U.S. west to Missouri, Oklahoma, and Texas). Bottomland
forests, mesic upland forests, sand savannas, sand prairies, banks of stream
and rivers, oxbows, sloughs, and swamps; also roadsides.
Morphological
variation in Q. pagoda was analyzed statistically by R. J. Jensen
(1989). It is very closely related to Q. falcata, which sometimes occurs
in slightly drier habitats in the same region, and the two have sometimes been
considered varieties of a single species (Steyermark, 1963). Hybrids involving Q.
pagoda are rare in Missouri, with only the hybrid involving Q. velutina
documented thus far.