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Description:
Dioecious trees, 0.5–3.0 m tall, monocaulous to sparsely branching. Leaves once-pinnate, 5.5–15 cm long; leaflets 17-21, lamina medium to green, dull above, paler below, coriaceous, lanceolate to ovate (terminal leaflet often obovate), 0.7–2.9 × (3–) 7–14 cm, the apex rounded to acute to acuminate, margin sharply spinulose dentate, thickened and revolute, the base slightly asymmetric, acute to rounded; petiolules 0–2.5 mm long; rachis articulated at insertion of leaflets, glabrous; petiole <5 mm (lowest pair of leaflets usually attached at base of leaf), 1.5–3 mm diam., glabrous, with a slightly alate base. Inflorescence an umbel or panicle of umbellules, terminal, erect, rhomboidal to obconic, the axes glabrous throughout; the bractlets often persistent, narrowly triangular (to linear), 2–4 mm long; the primary axis 2.7–5.2 (–10.5) cm long; the secondary axes 3–12, scattered or umbellate, 1.5–6.0 cm long; the tertiary axes (3–) 6–10 per secondary axis, 7–22 mm long, each with a terminal umbellule of 2–3 (–4) flowers (and rarely 1 lateral flower), flowers subtended by a linear to narrowly triangular bractlet 1.0–2.0 mm long; pedicels (1.0–) 2.0–5.0 mm long and 0.5–0.8 mm diam. in pistillate flowers, 1.0–2.0 mm long and 0.2–0.5 mm diam. in staminate flowers, articulated below the ovary. Calyx collar-like, ca. 0.2–0.5 mm tall, with 5 apiculate to narrowly triangular teeth. Corolla in bud oblong to conic in pistillate flowers, oblong (with minutely reflexed tips) in staminate flowers; the petals 5 in pistillate flowers, 4 in staminate flowers, not seen at anthesis. Stamens not seen at anthesis. Ovary 2-carpellate, elliptic to obovate, 6–7 mm tall at anthesis in pistillate flowers; ovary of staminate flowers vestigial, cylindrical; styles 2, free to fuse basally (but no more than 1/5 their lenghts), 1.2–1.6 mm long with the arms erect to ascending when receptive and spreading to recurved in fruit. Mature fruit compressed laterally, yellow-green with the apex deep red, then turning purple, elliptic to obovate, (5–) 7–8 × 4.5–5.0 mm, glabrous, the base rounded, the ribs weakly developed when dry.
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