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Region Country
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Specimen coordinates in square brackets [ ] have been approximated based on political units.
Country UpperLowerElevationLatitudeLongitudeDateQual.CollectorsColl NoInstitutions
BoliviaLa PazFranz Tamayo 800 m 14°22'00"S 068°29'00"W 7 Dec 1993 M. Kessler3891LPB
BoliviaSanta CruzVelasco 180 m 14°43'10"S 061°09'34"W 12 May 1995 René Guillén V. & R. Choré3588GOET, K, MO, NY, UC, USZ
BurundiBururi 800 m 22 Dec 1969 J. Lewalle4239 
BurundiMuramvya 1900m 23 janvier 1966 J. Lewalle327MO
CameroonAdamaoua 1634 m 07°22'09"N 013°59'17"E 3 octobre 2011 Vincent Droissart, G. Kamdem, Narcisse Kamdem & Pierre Ploton1083BRLU, MO, YA
CameroonCentral 1160 m 03°55'25"N 011°22'05"E 12 mai 2006 Vincent Droissart39BRLU
CameroonCentral 695 m 03°25'03"N 011°29'13"E 6 octobre 2013 Vincent Deblauwe70BRLU
CameroonCentral 903 m 03°58'18"N 011°24'36"E 19 septembre 2013 Vincent Deblauwe42BRLU, P
CameroonLittoral 1951 m 05°03'01"N 009°50'39"E 13 juillet 2014 Vincent Deblauwe & Rüksan Bose112BRLU, MO, P, YA
CameroonSouth 993 m 03°14'47"N 010°33'47"E 9 juin 2015 Vincent Deblauwe & Vincent Droissart233BRLU, MO, P, YA
CameroonSouthwest 1750 m 05°04'00"N 009°44'00"E 6 October 1986 M. Etuge & Duncan W. Thomas319MO
CameroonSouthwest 900 m 04°04'17"N 009°08'30"E 16 octobre 2013 Vincent Droissart, G. Kamdem, Moses Libalah, Thomas Couvreur, Sauquet, Hervé, J. Massoni & C. Wonkam1538BRLU, MO, YA
CameroonSouthwest 400 m 04°04'23"N 009°04'55"E 18 octobre 2013 Vincent Droissart, G. Kamdem, Moses Libalah, Thomas Couvreur, Sauquet, Hervé, J. Massoni & C. Wonkam1566BRLU, MO, YA
CameroonSouthwest 300 - 600 m 04°05'00"N 009°05'00"E 24 April 1984 Duncan W. Thomas3462MO
CameroonSouthwest 50 m 04°55'00"N 008°50'00"E 06 July 1983 - 16 July 1983 Duncan W. Thomas2227MO
CameroonSouthwest 500 m 04°05'00"N 009°06'00"E 29 December 1983 Duncan W. Thomas2822MO
CameroonSouthwest 200 - 300 m 04°38'00"N 009°26'00"E 18 October 1986 Stephen D. Manning508MO
Central African RepublicBamingui-Bangoran 585 m 08°30'00"N 021°13'00"E 20 November 1983 J.M. Fay6259MO
Central African RepublicBamingui-Bangoran 540 m 08°32'00"N 021°12'00"E 5 Jan 1984 J.M. Fay6306MO
Central African RepublicBamingui-Bangoran 590 m 16 Aug 1983 J.M. Fay5557MO
Central African RepublicVakaga 1330 m 09°10'00"N 023°13'00"E 01 November 1989 David J. Harris & J.M. Fay2133MO
ColombiaAntioquia 960 m 06°40'00"N 076°26'00"W 21 September 1987 James L. Zarucchi, Alan E. Brant & et al.5530HUA, MO
ComorosGrande Comore (Ngazidja) 7 August 1987 William G. D'Arcy17541MO
Costa RicaSan JoséAcosta 1460 m 09°44'54"N 084°11'02"W 30 July 2007 Michael H. Grayum, Barry E. Hammel, J. F. Morales, Isabel Pérez & Lady Muñoz12493MO
Costa RicaSan JoséDota 1720 m 09°32'05"N 083°50'19"W 26 agosto 2004 Alexander Rojas & Hernán Gómez5984MO
Costa RicaSan JoséEscazu 1850 m 09°52'40"N 084°08'13"W 09 septiembre 2004 Alexander Rojas & Rafael Chacón6026MO
Democratic Republic of the CongoKatanga 1250 m 11°10'48"S 027°05'24"E 03 April 1990 Robert R. Brooks, A. Baker, Henk Schat & François Malaisse78MO
Democratic Republic of the CongoManiema 470 m 02°42'16"S 025°07'39"E 10 avril 2015 Roy E. Gereau, Gilbert Paluku, Jeanne Mbey & P. Akimo7474BR, MA, MO, WAG
EcuadorCotopaxi 950 - 1000 m 00°53'00"S 079°10'00"W 06 Apr 1973 L.B. Holm-Nielsen, S. Jeppesen, B. Løjtnant & Benjamin Øllgaard3103AAU, MO
EcuadorGalapagos 470 m 13 Feb 1976 S. Jeppesen3044AAU
EthiopiaGamo Gofa 1550 m 06°17'00"N 036°18'00"E 8 November 1984 Roy E. Gereau1405MO
Gabon [00°48'39"S] [011°51'28"E] 1990 J.J. Dibata666 
GabonNgounie 01°10'00"S 011°49'00"E 14 February 1988 Adriaan M. Louis2768MO, UC
GabonNgounie 125 m 02°17'43"S 011°17'52"E 9 décembre 2016 Ehoarn Bidault, Archange Boupoya & Elie Nzigou Doubindou2968BRLU, LBV, MO, P
GabonNgounie 146 m 01°23'16"S 010°52'57"E 12 November 2009 Tariq Stévart, D. Ngema, Etienne Mounoumoulossi, P. Bissiemou & Eric Akouangou3327LBV, MO
GhanaEastern 810 m 06°13'48"N 000°32'42"W 4 July 1995 Daniel K. Harder & et al.3344 
GhanaEastern 500 - 700 m 06°09'47"N 000°36'15"W 14 November 1995 Heidi H. Schmidt & et al.1679 
GuineaConakry 946 m 07°40'30"N 008°23'42"W 22 June 2012 Tariq Stévart, Jean-Yves Serein & Gononan Traoré4545MO, P, SERG, WAG
GuineaConakry 728 m 07°38'48"N 008°21'20"W 26 June 2012 Tariq Stévart & Jean-Yves Serein4560BRLU, MO, P, SERG, WAG
GuineaNzérékoréLola 1476 m 07°39'14"N 008°22'35"W 19 July 2008 C. C. H. Jongkind8242WAG
GuineaNzérékoréLola 1675 m 07°39'55"N 008°22'46"W 17 September 2011 C. C. H. Jongkind, Moussa Diabaté, Peter B. Phillipson & Gordon McPherson11112BR, G, K, MO, P, SERG, WAG
GuineaNzérékoréLola 1300 m 07°40'33"N 008°22'45"W 05 octobre 2011 D. Bilivogui, Moussa Diabaté, Tariq Stévart & Ehoarn Bidault70BR, MO, P, SERG, WAG
GuineaNzérékoréLola 1139 m 07°41'03"N 008°22'29"W 31 octobre 2012 D. Bilivogui & Elodie Lerat214BR, MO, P, SERG, WAG
GuineaNzérékoréLola 1020 m 07°40'36"N 008°23'18"W 02 October 2011 Gordon McPherson21509MO
GuineaNzérékoréLola 1050 m 07°40'38"N 008°23'15"W 07 octobre 2011 Moussa Diabaté, D. Bilivogui, Gordon McPherson & Ehoarn Bidault983MO, P, SERG
GuineaNzérékoréLola 1000 m 07°40'44"N 008°23'08"W 19 September 2011 Gordon McPherson21346MO
KenyaWesternKakamega 1540 m 00°10'12"N 034°31'12"E 15 October 2008 P. Kamau94EA, MO
LiberiaLofa 518 m 08°07'42"N 009°58'59"W 19 October 2019 Philomena Yarwoah, Ehoarn Bidault, Kolou Koïvogui & Bruno Senterre35BRLU, K, MO, P, SERG
Madagascar [18°56'47"S] [047°06'03"E]   Georges Cremers3699MO, TAN
Madagascar [18°56'47"S] [047°06'03"E]   Jean M. Bosser12568MO, TAN
Madagascar [18°56'47"S] [047°06'03"E]   Jean M. Bosser8053MO, TAN
MadagascarAntananarivo 1200 m 18°19'00"S 045°34'00"E 23 January 1985 L.C. Barnett & et al.392MO
MadagascarAntananarivoAnalamanga 18°09'00"S 047°18'00"E   des Abbayes2673 
MadagascarAntananarivoAnalamanga 18°45'00"S 047°34'00"E   A. C. D'Alleizette172 
MadagascarAntananarivoItasy 891 m 18°59'37"S 046°29'59"E 27 janvier 2015 Richard Razakamalala, N. Manjato & J. Razafimandimby7891MO, P, TAN
MadagascarAntananarivoVakinankaratra 1470 m 20°03'49"S 047°00'02"E 09 February 2018 Ulf Swenson, V. Razafindrahaja, Anselme Atalahy & Sylvain G. Razafimandimbison1893MEL, MO, P, S, TAN
MadagascarAntsiranana 1567 m 14°10'47"S 049°22'01"E 20 February 2007 Désiré Ravelonarivo, Charles Rakotovao & Richard Razakamalala2199G, MO, P, TAN
MadagascarAntsirananaDIANA 600 - 900 m 14°05'00"S 048°23'00"E 26-30 March 1993 Simon T. Malcomber, Pierre J. Rakotomalaza & Jeannine I. Raharilala2284MO
MadagascarAntsirananaDIANA 1000 - 1480 m 14°04'00"S 048°17'00"E April 1993 Pierre J. Rakotomalaza32 
MadagascarAntsirananaDIANA 600 - 900 m 14°05'00"S 048°23'00"E 26-30 March 1993 Simon T. Malcomber, Pierre J. Rakotomalaza & Jeannine I. Raharilala2312MO
MadagascarAntsirananaDIANA 1100 m 14°02'50"S 048°47'09"E 26 Nov 2000 A. Rasolohery56MO, P, TAN, TEF
MadagascarAntsirananaDIANA 12°36'00"S 049°09'00"E   J.M. Hildebrandt3385 
MadagascarAntsirananaDIANA 13°17'00"S 048°16'00"E   L.H. Boivin218 
MadagascarAntsirananaDIANA 1470 m 12°35'47"S 049°09'08"E 14 novembre 2007 L. Nusbaumer2469G, K, MO, P, TEF, WAG
MadagascarAntsirananaDIANA 1042 m 12°34'46"S 049°07'37"E 23 janvier 2012 S.D. Ramandimbimanana & C.H. Randimbiarison280G, MO, P, TEF
MadagascarAntsirananaDIANA 1120 m 12°35'59"S 049°10'58"E 30 avril 2008 M.H. Razanajatovo, S.D. Ramandimbimanana & S.M. Trigui123G, K, MO, P, TEF
MadagascarAntsirananaDIANA 1065 m 12°34'52"S 049°08'10"E 13 janvier 2008 S.M. Trigui, M.H. Razanajatovo & S.D. Ramandimbimanana87G, K, MO, P, TEF, WAG
MadagascarAntsirananaDIANA 1150 m 12°37'02"S 049°10'20"E 11 avril 2008 S.M. Trigui, M.H. Razanajatovo & S.D. Ramandimbimanana241G, TEF
MadagascarAntsirananaDIANA 980 m 12°36'15"S 049°11'44"E 28 avril 2008 S.M. Trigui, M.H. Razanajatovo & S.D. Ramandimbimanana248G, K, MO, P, TEF, WAG
MadagascarAntsirananaSAVA 1004 m 14°20'12"S 049°17'09"E 11 November 2006 Charles Rakotovao & et al.3316 
MadagascarAntsirananaSAVA 860 m 13°17'31"S 049°35'35"E 17 novembre 2005 L. Nusbaumer & P. Ranirison1622G
MadagascarAntsirananaSAVA 811 m 14°10'24"S 049°18'21"E 06 Feb 2006 Richard Razakamalala & et al.3098MO, P, TAN
MadagascarAntsirananaSAVA 811 m 14°10'24"S 049°18'21"E 06 Feb 2006 Richard Razakamalala & et al.3110MO, P, TAN
MadagascarAntsirananaSAVA 910 m 14°10'22"S 049°17'54"E 06 Feb 2006 Richard Razakamalala & et al.3176MO, P, TAN
MadagascarAntsirananaSAVA 957 m 14°09'46"S 049°22'34"E 25 Feb 2007 Richard Razakamalala & et al.3334MO, P, TAN
MadagascarAntsirananaSAVA 738 m 14°21'34"S 049°22'35"E 05 May 2006 Charles Rakotovao & et al.3149MO, P, TAN
MadagascarFianarantsoaAmoron'i Mania 1488 m 20°28'40"S 046°33'34"E 25 April 2003 Tefy Andriamihajarivo191MO, P, TAN
MadagascarFianarantsoaAmoron'i Mania 20°33'00"S 046°48'00"E   Raymond Decary13127 
MadagascarFianarantsoaAmoron'i Mania 1463 m 20°25'23"S 047°10'09"E 12 March 2010 Jacky Andriantiana, Fano A. Rajaonary, M. Rabarimanarivo, A. Ramahefaharivelo, Nivo H. Rakotoarivelo, Hanta Razafindraibe, C. Frasier, Brigitte Ramandimbisoa, S. Porembski, Rakotonandrasana & Randriamandimbiarisoa675MO, P, TAN
MadagascarFianarantsoaAmoron'i Mania 1482 m 20°25'12"S 047°09'59"E 12 March 2010 Jacky Andriantiana, Fano A. Rajaonary, M. Rabarimanarivo, A. Ramahefaharivelo, Nivo H. Rakotoarivelo, Hanta Razafindraibe, C. Frasier, Brigitte Ramandimbisoa, S. Porembski, Rakotonandrasana & Randriamandimbiarisoa707MO, P, TAN
MadagascarFianarantsoaMatsiatra Ambony (formerly Haute Matsiatra) 1200 m 21°31'52"S 047°01'32"E 14 March 2010 Brigitte Ramandimbisoa, A. Ramahefaharivelo, C. Frasier, Fano A. Rajaonary, M. Rabarimanarivo, Hanta Razafindraibe, Jacky Andriantiana, Nivo H. Rakotoarivelo, Rakotomalala, Jarison & Randrianantenaina, J.C. Emile8MO, P, TAN, TEF
MadagascarFianarantsoaMatsiatra Ambony (formerly Haute Matsiatra) 1327 m 21°07'38"S 047°10'46"E 13 March 2010 M. Rabarimanarivo, Jacky Andriantiana, Fano A. Rajaonary, Nivo H. Rakotoarivelo, N. Ravalimanarivo & Voahanginiaina, Ernestine179MO, P, TAN
MadagascarFianarantsoaMatsiatra Ambony (formerly Haute Matsiatra) 1158 m 21°52'56"S 046°49'18"E 17 March 2010 A. Ramahefaharivelo, M. Rabarimanarivo, Fano A. Rajaonary, Nivo H. Rakotoarivelo, Brigitte Ramandimbisoa, Hanta Razafindraibe, N. Ravalimanarivo & Ratsimbazafy, Landri82MO, P, TAN
MadagascarMahajanga 30 mars 1993 J.F. Villiers, Jens Klackenberg & F. Badré5001MO, P
MadagascarMahajanga 3910 ft 15°31'10"S 049°05'40"E 25 Apr 2007 Désiré Ravelonarivo2353MO, P, TAN
MadagascarMahajangaBetsiboka 17°36'00"S 046°55'00"E   H. Perrier de la Bâthie468 
MadagascarMahajangaBetsiboka 16°56'00"S 046°57'00"E   Raymond Decary19260 
MadagascarMahajangaBoeny 16°58'00"S 044°24'00"E   H. Perrier de la Bâthie7436 
MadagascarMahajangaMelaky 220 m 17°54'36"S 044°28'44"E 8 février 2012 R. Bolliger, R. M. Hanitrarivo & B. F. L. Rakotozafy188BR, G, K, MO, P, TEF
MadagascarMahajangaMelaky 297 m 18°03'36"S 044°31'09"E 28 février 2012 R. M. Hanitrarivo, R. Bolliger & B. F. L. Rakotozafy233BR, G, K, MO, P, TEF
MadagascarMahajangaMelaky 18°40'00"S 044°42'00"E 25 mars 1993 J.F. Villiers, Jens Klackenberg & F. Badré4871MO, P
MadagascarMahajangaSofia 840 m 14°47'49"S 048°23'47"E 29 février 2020 Richard Razakamalala8617MO, P, TAN
MadagascarToamasinaAlaotra-Mangoro 1117 m 18°51'37"S 048°17'55"E 02 Mar 2005 P. Antilahimena3537MO, P, TAN
MadagascarToamasinaAlaotra-Mangoro 1140 m 18°48'27"S 048°19'00"E 15 dec 2004 Lalao Andriamahefarivo & et al.15MO, P, TAN
MadagascarToamasinaAlaotra-Mangoro 1025-1100 m 17°44'51"S 048°42'47"E 01 Mars 2001 F. Ratovoson436MO
MadagascarToamasinaAlaotra-Mangoro 1070 m 18°48'41"S 048°20'08"E 17 December 2005 P. Antilahimena, Félix Edmond & Tojo4424AG, MO, P, TAN, UC
MadagascarToamasinaAlaotra-Mangoro 1083 m 18°48'34"S 048°19'21"E 17 Feb 2005 P. Antilahimena & et al.3393MO, P, TAN
MadagascarToamasinaAlaotra-Mangoro 1124 m 18°48'47"S 048°19'21"E 24 Feb 2005 P. Antilahimena & et al.3475MO, P, TAN
MadagascarToamasinaAlaotra-Mangoro 1127 m 18°48'33"S 048°19'39"E 25 Feb 2005 Charles Rakotovao & et al.1377MO, P, TAN
MadagascarToamasinaAlaotra-Mangoro 1078 m 18°48'24"S 048°20'19"E 4 avril 2005 Charles Rakotovao & Félix Edmond1806MO, P, TAN
 
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