4. Solidago caesia L. (blue-stemmed goldenrod, wreath goldenrod)
Pl. 239 c–e; Map
993
Plants with the
rootstock short, stout, and sometimes branched, sometimes also producing long,
slender rhizomes. Stems 1 to several, 30–100 cm long, erect to more commonly
arched or pendant, sometimes with fine, inconspicuous, longitudinal lines but
not noticeably ridged or grooved, glabrous, not shiny, glaucous. Leaves chiefly
cauline, the largest leaves in the lower 1/3 of the stem, the basal and lower
stem leaves absent at flowering. Basal and lowermost stem leaves with the blade
6–10 cm long, 1–3 cm wide, mostly 3–8 times as long as wide, narrowly
elliptic-oblanceolate to elliptic or elliptic-obovate, relatively thin, tapered
to a sessile base, tapered to a sharply pointed tip, the margins sharply
toothed and usually inconspicuously hairy, the surfaces glabrous, the
undersurface with 1 main vein, the fine, pinnate secondary veins usually
relatively easily observed (these usually forming an irregular network). Median
and upper stem leaves 2–8 cm long, elliptic-oblanceolate to narrowly elliptic,
the margins of the uppermost leaves sometimes entire, otherwise similar to the
lower stem leaves. Inflorescences of axillary clusters, the heads oriented in
several directions. Involucre 2.5–4.5 mm long, the bracts in 3–5 unequal
series. Involucral bracts mostly oblong to narrowly oblong and rounded to
bluntly pointed (those of the outer series often oblong-lanceolate and sharply
pointed) at the appressed-ascending tip, the thin, white to yellowish white
margins hairy toward the tip, the outer surface glabrous, with a poorly
defined, green central region toward the tip, this tapered abruptly to the
midvein above or below the bract midpoint, the midvein often slightly thickened
and sometimes with a faint, additional pair of veins present. Receptacle naked.
Ray florets (1–)2–5, the corollas 3.0–3.5 mm long, yellow. Disc florets 5–9,
the corollas 3.0–3.5 mm long, the lobes 0.9–1.5 mm long, yellow. Pappus 2.5–3.0
mm long, a few of the bristles often slightly thickened toward the tip. Fruits
1.0–1.8 mm long, narrowly ellipsoid-obovoid, finely hairy. 2n=18. August–October.
Scattered in the
southern quarter of the state, almost entirely within the Ozark Division
(eastern U.S. west to Wisconsin and Texas; Canada). Bases and ledges of shaded
bluffs, banks of streams and rivers, bottomland forests, and mesic upland
forests; also roadsides.
This species is
found most frequently along sheltered bluffs and adjacent lower terraces of
streams. It does well in full sun or partial shade in the wildflower garden
with its graceful, arching, glaucous stems and attractive clusters of heads.
Cook and Semple
(2004) recently divided the species into two varieties, based mainly on
differences in leaf morphology. They segregated plants with broadly lanceolate
to elliptic or elliptic-ovate median leaves 5–9 cm long and less-arched stems
as var. zedia R.E. Cook & Semple, which they stated to occur from
southern Arkansas through Mississippi into northern Florida. They characterized
the var. caesia as having narrowly lanceolate midstem leaves 5–15 cm
long and strongly arched stems. Although Cook and Semple did not include
Missouri in the range of var. zedia, Rachel Cook annotated about a third
of the specimens from throughout the species range in the state as this variety.
The determinations of these specimens seem equivocal, with a number of the
Missouri specimens seemingly intermediate in leaf shape. None of the specimens
in question include observations by the collectors on stem arching. In light of
the difficulties in applying the defining features of these infraspecific taxa
to plants in the Missouri flora, it seems wisest for the present time not to
attempt a formal recognition of varieties.