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(Last Modified On 5/8/2013)
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(Last Modified On 5/8/2013)
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Species
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Gouania lupuloides (L.) Urban
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PlaceOfPublication
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Symb. Antill. 4: 378. 1910.-FIG. 8.
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Synonym
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Banisteria lupuloides L., Sp. P1. 427. 1753. Rhamnus polygamus Jacq., Enum. P1. Carib. 17. 1760. Gouania polygama (Jacq.) Urban, Symb. Antill. 4: 378. 1910.
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Description
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Lianas, or scandent to arching shrubs, the tendrils at the bases of the inflorescences, the younger stems pubescent, the trichomes grey to brown. Leaves inconspicuously 3-nerved at the base, ovate or sometimes elliptic, acute, serrate, the teeth mostly glandular and remote, rounded to shallowly cordate basally, to 11(-13) cm long and 7(-9) cm wide, glabrous to sparsely pubescent above, more or less glabrous, appressed-pubescent on the veins to densely pubescent beneath; petioles 0.5-1.5(-2) cm long, pubescent, the stipules minute, deciduous.
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Habit
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Lianas shrubs
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Description
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Inflorescences glomerules, sessile or subsessile, 5-8-flowered, arranged in "spikes" 8-18 cm long, each glomerule subtended by a bract, deltoid, 2-3 mm long, pubescent, the rachis pubescent. Flowers mostly bisexual, sometimes functionally carpellate or staminate by reduction; subsessile or with pedicels 1-2 mm long; floral tube more or less obconic, pubescent; sepals deltoid, ca. 0.6-1 mm long, keeled on the inner surface, densely pubescent on the outer surface; petals white, ca. 0.8-0.9 mm long; stamens functional or reduced, the filaments ca. 0.5-0.8 mm long, the anthers ca. 0.15-0.2 mm long; nectariferous disc 1- or 2-lobed between the stamens, generally 10-ridged, sparsely pubescent near the emergence of the styles; ovary functional or sometimes reduced, the styles 3, ca. 0.6-0.9 mm long. Schizocarps sparsely pubescent on the body, rarely densely so, the body brown to green-black, ca. 3-4 mm high, the wings paler, dolabriform, ca. 5-6 mm high and 4-5 mm wide; seeds 3, brown, ca. 2 mm. long.
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Distribution
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A variable and wide-ranging species found in Mexico, Central America, the West Indies, and northern South America.
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Specimen
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BOCAS DEL TORO: Vicinity of Chiriqui Lagoon, von Wedel 1834, 1850, 1898 (all MO), 2045 (GH, US). Farm 8, Cooper 162 (F, GH, US). CANAL ZONE: Balboa, Standley 25531, 32137 (both US). Barro Colorado Island, Croat 4709, 7984 (both MO), Shattuck 444 (MO), This content downloaded from 192.104.39.2 on Tue, 7 May 2013 17:30:03 PMAll use subject to JSTOR Terms and Conditions1971] NOWICKE-FLORA OF PANAMA (Family 111. Rhamnaceae) 283 523 (F), Wetmore & Abbe 48, 67 (both A, GH), Woodworth & Vestal 326 (A, F), 351 (A). Chagres, Fendler 108 (GH, US). Along dirt road to Chiva-Chiva, Correa A. 498 (MO). 5 mi. N of Cocli, Tyson 3873 (MO). Darien Station, Standley 31604 (US). Cocoll Road, Burch et al. 1395 (MO). Between France Field and Catival, Standley 30249, 30372 (both US). Las Cruces Trail, Hunter & Allen 702 (MO), Standley 29202 (US.). Vicinity of Miraflores, White & White 53 (A, F). Along Pan American Highway, Nowicke et al. 3580 (MO). COCLE: Aguadulce, Pittier 4903 (US). 4 mi. W of Anton on Rio Chico, Tyson & Blum 2590 (MO). El Valle de Anton, Allen 2863 (F, MO), Harvey 5165 (F). DARIEN: Trail between Cana and Boca de Cupe, Stern et al. 625 (GH, MO, US). Vicinity of Pinogana, Allen 4290 (F, MO). Along Rio Pinas, Duke 9274, 10550 (both MO). Along Rio Sambu', Pittier 5561 (US). Tucuti, Terry & Terry 1378, 1384 (both A, F, MO). HERRERA: 4 mi. S of Los Pozos, Tyson 2674 (MO). 0.5 mi. N of Macaracas of Rio La Villa, Tyson et al. 3138 (MO). Vicinity of Ocu', Stern et al. 1715 (MO, US). LOSi SANTOS: Punta Mala near mouth of Rio Caldera, Tyson 2733 (MO.). PANAMA: Vicinity of Bella Vista, Piper 5346 (GH, US), Standley 25398 (MO, US.). Chepillo Island, Duke 10316 (MO). Sabanas near Chepo, Hunter & Allen 99 (MO). Between Matias Hernandez and Juan Diaz, Standley 28962 (A, US) 31999 (US). Sabanas N of Panama City, Bro. Paul 597 (US). Punta Paitilla, Piper 5397 (US ). San Jose Island, Erlanson 18 (GH, US.), Johnston 514 (GH, MO), 619 (GH), 906, 1173 (both GH, MO, US), Tyson & Loftin 5083 (MO). Taboga Island, Standley 27051 (MO, US), 27906 (US). Taboguilla Island, Miller 1989 (MO, US). Swamp E of Rio Tocumen, Standley 26557 (US.). VERAGUAS: Vicinity of Santiago, Allen 1078 (F, GH, MO, US). PROVINCE UNKNOWN: Camino de Corozal, Bro. Heriberto 250 (GH, US).
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Note
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One collection has distinctively large fruits, a characteristic also found in a Mexican species, G. conzattii Greenman. However, comparison with the type of G. conzattii (Conzatti 1567, F) indicates two different taxa. The material is insufficient for precise identification: Darien: Vicinity of Cana, Williams 723 (NY, US). Leaves ovate, acute, remotely crenate, the teeth glandular-tipped, shallowly cordate basally, 10-11 cm long and 6-7 cm wide, more or less glabrous on both surfaces, slightly coriaceous; petioles ca. 1.5 cm long, more or less glabrous, the stipules apparently deciduous. Inflorescences arranged in "spikes" to 18 cm long, the rachis pubescent. Flowers unknown. Schizocarp pubescent, the body and wings pale brown, the body 7-8 mm high, the wings reniform, 10-11 mm high and 6-7 mm wide; seeds 3, wedge-shaped, ca. 5 mm long.
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