Liana; young branches solid or hollow in the center, indumentum dense to sparse pilose or sericeous (rarely tomentose), white and/or light red-brown. Leaf blades 4—13.2 x 2—8 (9.2) cm, narrow to wide elliptic to ovate or lanceolate, membranaceous, both surface drying similar green, less frequently brown-green (rarely red-brown) or abaxial surface drying slightly paler, lateral veins (3) 4 or 5, less frequently 6, pairs, at 45—60 (70)º angle to midrib, adaxial surface abundant pubescent to glabrescent, the trichomes 0.2—1 mm, appressed, ascending, straight to slightly curved, sometimes some or all spreading, or torulose but still appressed, white or yellow-white (rarely red-brown), abaxial surface dense pubescent to glabrescent, the trichomes 0.15--1 mm, sometimes only on venation, appressed, straight and ascending, or some or all somewhat curving or torulose or straight and spreading in all directions including erect, overall or only on veins or vein axils, white or yellow-white (rarely red-brown), base subcordate, rounded or cordate and then usually shortly decurrent, margin serrate to serrulate to nearly entire, irregularly serrate or 1—3 (4) teeth/cm, tooth glands cupular (rarely pulvinate), apex acuminate or cuspidate (rarely acute); stipules 2-lobed,usually retained (at least in part) into fruiting, upper lobe 1--10 x 0.5--3 mm, curving upward or erect, lanceolate, acuminate and the tip flexuous or triangular, acuminate, firm,lower lobe 0.1--14 mm, curving downward or spreading, lanceolate-acuminate, linear, subulate, oblong, ovate, circular, broadly rectangular, or reniform, sometimes with long extended glandular-tip or with 2 or 3 short to long-extended glandular teeth;petioles 3--26 mm. Inflorescence with longest racemiform part 7--25 cm, indumentum of rachis sparse to dense pilose or tomentose, white or yellow-white, and/or red-brown, bract of cyme 1.2—4 (--7) mm, lanceolate, acuminate, cyme appearing sessile (rarely peduncle to 1.5 mm). Mature flowers with pedicels (0.5--) 1—3 (--4) mm,hypanthium sparse to dense sericeous (rarely tomentose), the trichomes white, yellow-white or red-brown. Sepals (0.5--) 0.75—1.6 mm, externally sparse to abundant sericeous, the trichomes white, yellow-white or red-brown. Petals (0.5--) 0.6—1.5 mm; stamens with filaments (0.25--) 0.3—1.2 mm, narrowly triangular to linear, anthers (0.15--) 0.2—0.3 mm; disc excluding lobes 1—1.8 (–2.1) mm diam., glabrous, disc lobes (0.1-- ) 0.2—0.5 x (0.1--) 0.2—0.75 mm at apex, square to transversely rectangular (rarely longer than wide and narrowing apically, very rarely to point), generally apically bi-lobed, 1/5—2/5 (1/2) length of sepals; style glabrous (rarely with trichomes).Young fruits glabrescent to pilose or sericeous with trichomes sparse, white to light red-brown; mature mericarp out-line butterfly shaped with emarginations at apex and base, the wings attached to apex and base of fruit body as well as sides, externally drying yellow-brown, green-brown, red-brown or yellow, darker brown over fruiting body, glabrous or fruit body apically sparse and minutely pilose, puberulent or sericeous with trichomes 0.1—0.3 mm, white, internally drying off-white on wings, slightly or moderately contrasting with slightly darker off-white to beige fruit body, fruit body 4--7 mm high, wings (5) 6.6--12 mm high, distance between highest points of two wings 3—7 mm, width of mericarp (8) 9--16 mm, 1.5—3.2 times height of fruit body, width of fruit body 3—4.5 (5) mm, (1/5) ¼--1/2 width of mericarp. Seeds 2.7—3.8 x (1.5) 2--3 mm, shiny brown.