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(Last Modified On 9/10/2013)
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(Last Modified On 9/10/2013)
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Species
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Joosia panamensis Dwyer
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Note
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TYPE: Panama, Croat 25932 (MO).
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Description
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Shrubs or small trees, the branchlets glabrescent, often covered with mosses. Leaves elliptic or rarely obovate trapeziform, 16-22 cm long, 6-9.5 cm wide, acute or widely deltoid toward the apex, cuneate or acute at the base, slightly inequilateral, the costa prominulous above, prominent beneath, the lateral veins 10-15, arcuate, the veinlets dense, arising at right angles from the costa, stiffly chartaceous, glabrescent above, beneath with white, appressed hairs; petioles to 1.5 cm long, villose; stipules free, ovate oblong, ca. 2 cm long, ca. 0.5 cm wide at the base, obtuse, red when dry, the hairs few, white. Inflorescences terminal, perhaps solitary, cymose umbellate, the peduncle, rigid, slender, to 8 cm long, ca. 0. 15 cm wide, often twisted, densely pubescent, the flowers secundly disposed and crowded on short branches arising radiately from the apex of the peduncle; pedicels ca. 2 mm long. Flowers with the calyx cup ca. 2.5 mm long, ca. 3 mm wide, cup shaped, expanded, densely ciliolate outside, glabrous within, the lobes 4, oblong, ca. 2 mm long, to 1.5 mm wide, rounded at the apex, glabrous within; corolla white, the tube narrowly cylindrical, ca. 9 mm long, ca. 1.8 mm wide, carnose, the hairs outside appressed, minute, glabrous within, the lobes 5, ovate oblong, to 3 mm long, with 2 ovate elliptic appendages within, to 2.8 mm long, petaloid, glabrous, scalloped; anthers 5, sessile or subsessile or filaments to 2.5 mm long, narrow oblong, ca. 3.5 mm long, attached to the tube at or below the middle, often near the base; style (2-)4-10 mm long, the stigmas 2, unequal, scarcely wider than the style, to 2 mm long. Fruits linear oblong, to 2 cm long, to 0.35 cm wide, terete, scarcely angular, the ribs few, prominulous, minutely glandular punctate; seeds spindle shaped ca. 6 mm long, the wing bifid or ero- sulose at the base, acute, entire or fimbriate at the apex.
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Habit
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Shrubs or small trees
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Distribution
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known only from Panama.
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Note
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It is related to J. umbellifera Karst., a well-known Colombian species. It differs in having much shorter fruits, which measure only about a 1/3 as long as those of J. umbellifera, much shorter seeds, and presumably shorter stipules.
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Specimen
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BOCAS DEL TORO: Chiriqui Trail between Buena Vista Coffee Finca & Cerro Pil6n, Kirkbride & Duke 680 (MO). COCLE: Alto Calvario around Rivera sawmill, 7 km N of El Cope, 700-900 m, Folsom 3240 (MO). COL6N: 2-3 mi up Rio Guanche, Kennedy & Foster 2136 (MO). PANAMA: 16-20 km above Panamerican Highway road from El Llano to Carti-Tupile, 400 m, Kennedy 2694 (MO). El Llano- Carti Road, 18 km from Panamerican Highway, Mori et al. 4592 (MO). VERAGUAS: Road between Escuela Agricola Alto de Piedra above Santa F6 and Rio Dos Bocas, 730-770 m, Croat 25932 (MO). Valley of Rio Dos Bocas between Alto Piedra and Calovebora, 350-400 m, Croat 27471 (MO). Caribbean slope above Rio Primero Brazo, S mi NW of Santa Fe, Liesner 1014 (MO). 8.8 km from Escuela Agricola Alto de Piedra, Mori & Kallunki 3220, 3239 (both MO). 16 km from Santa F6, near Rio Caloveborita, Mori 6688 (MO).
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