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  • Reference article Brako, L. & J. L. Zarucchi. (eds.) 1993. Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45: i–xl, 1–1286.
  • Reference article Forzza, R. C. 2010. Lista de espécies Flora do Brasil http://floradobrasil.jbrj.gov.br/2010. Jardim Botânico do Rio de Janeiro, Rio de Janeiro.
  • Reference article Macbride, J. F. 1936. Rubiaceae, Flora of Peru. Publ. Field Mus. Nat. Hist., Bot. Ser. 13(6/1): 3–261. View in Biodiversity Heritage Library
  • Reference article Neill, D. A. & C. Ulloa Ulloa. 2011. Adiciones Fl. Ecuador: Segundo Supl., 2005-2010 1–202. Fundación Jatun Sacha, Quito.
  • Reference article Taylor, C. M. 2010-2025. Studies in Tropical Rubiaceae.
  • Reference article Taylor, C. M. & D. Zappi. 2006. Rubiaceae (pt. 5), Tribe 18. Psychotrieae (2). Fl. Ecuador 79: 1–111.
  • Reference article Ulloa Ulloa, C., P. Acevedo-Rodríguez, S. G. Beck, M. J. Belgrano, R. Bernal, P. E. Berry, L. Brako, M. Celis, G. Davidse, S. R. Gradstein, O. Hokche, B. León, S. León-Yánez, R. E. Magill, D. A. Neill, M. H. Nee, P. H. Raven, H. Stimmel, M. T. Strong, J. L. Villaseñor Ríos, J. L. Zarucchi, F. O. Zuloaga & P. M. Jørgensen. 2017. An integrated assessment of vascular plants species of the Americas. Science 358: 1614–1617 [Online Suppl. Materials: 1–23 + 1–2497], f. 1–4 [f. S1–5].
  • Reference article Ulloa Ulloa, C., P. Acevedo-Rodríguez, S. G. Beck, M. J. Belgrano, R. Bernal, P. E. Berry, L. Brako, M. Celis, G. Davidse, S. R. Gradstein, O. Hokche, B. León, S. León-Yánez, R. E. Magill, D. A. Neill, M. H. Nee, P. H. Raven, H. Stimmel, M. T. Strong, J. L. Villaseñor Ríos, J. L. Zarucchi, F. O. Zuloaga & P. M. Jørgensen. 2018 [Onwards]. An integrated Assessment of Vascular Plants Species of the Americas (Online Updates).
 
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