Accepted By:
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AFPD. 2008. African Flowering Plants Database - Base de Donnees des Plantes a Fleurs D'Afrique.
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Bridson, D.M. & B. Verdcourt. 1988. Rubiaceae (part 2). 415–747. In W. B. Turrill & R. M. Polhill (ed.) Fl. Trop. E. Africa. A. A. Balkema, Rotterdam.
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Gibbs Russell, G. E., W. G. M. Welman, E. Retief, K. L. Immelman, G. Germishuizen, B. J. Pienaar, M. Van Wyk & A. Nicholas. 1987. List of species of southern African plants. Mem. Bot. Surv. South Africa 2(1–2): 1–152(pt. 1), 1–270(pt. 2).
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Schatz, G. E., S. Andriambololonera, P. P. Lowry II, P. B. Phillipson, M. Rabarimanarivo, J. I. Raharilala, F. A. Rajaonary, N. Rakotonirina, R. H. Ramananjanahary, B. Ramandimbisoa, A. Randrianasolo, N. Ravololomanana, C. M. Taylor & J. C. Brinda. 2023. Catalogue of the Plants of Madagascar.
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Taylor, C. M. 2010-2025. Studies in Tropical Rubiaceae.
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