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Published In: Species Muscorum Frondosorum 51–56, pl. 8. 1801. (Sp. Musc. Frond.) Name publication detailView in BotanicusView in Biodiversity Heritage Library
 

Project Name Data (Last Modified On 2/18/2011)
Acceptance : Accepted
Project data     (Last Modified On 2/18/2011)
Discussion:

Splachnum is predominantly a circumpolar, arctic-boreal genus (Koponen 1983) of 11 species (Crosby et al. 2000). Of these only S. weberbaueri is confined to the neotropics, while the eastern North American S. pensylvanicum (Brid.) Crum is disjunct in Venezuela (Steyermark 103922 MO). Although Splachnum has a single peristome, it is the only genus in the Splachnaceae that has not lost its endostome. The endostome is fully fused to the exostome teeth which are united into 8 pairs. Each tooth-pair has four columns of cell remnants on its outer surface as well as four columns on its inner surface.

Splachnum is an entomophilous genus with prominent turbinate capsule hypophyses and the odd feature that the setae continue to elongate after capsule maturation. Marino (1988) considered Splachnum and Tetraplodon unique in their restriction to organic substrates.


 

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Splachnum L. ex Hedw., Sp. Musc. Frond. 51. 1801. 

Plants soft, densely caespitose, pale yellowish green, somewhat shiny. Stems erect, forked below; rhizoidal tomentum dense, branched, reddish brown, cells smooth. Leaves erect to erect-spreading when wet, imbricate in lower half and variously contorted in upper half when dry; apex acuminate; margins plane below, somewhat inrolled above, indistinctly bordered at base by thin-walled, elongate cells; costa single, percurrent or slightly excurrent; upper cells large, pale, lax, thin-walled, irregularly rhombic, basal cells rectangular. Autoicous or dioicous. Perigonia terminal or axillary; perichaetia terminal. Setae elongate, smooth, twisted when dry. Capsules erect, firm, subcylindric, slightly contracted below the suboral rim; hypophysis inflated, turbinate, brightly colored, delicate; exothecial cells quadrate, with thickened, yellowish walls, suboral rim with thick-walled, oblate cells; opercula flat-convex, with a small apiculus; columella slightly protruding from the mature capsule mouth; peristome teeth 8, densely papillose, dark-red, incurved or erect when moist, tightly reflexed when mature and dry. Spores ellipsoidal, smooth, brown in mass. Calyptrae conic-mitrate, smooth, sometimes split on one side.

 
 
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