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Published In: Die Laubmoose Deutschlands, Oesterreichs und der Schweiz 1: 590. 1888. (Laubm. Deutschl.) Name publication detailView in Biodiversity Heritage Library

Project Name Data (Last Modified On 2/16/2011)
Acceptance : Accepted
Project data     (Last Modified On 2/16/2011)

Timmiella has bistratose leaves in which the ventral layer is bulging-mammillose and the dorsal layer plane. The cell layers near the costa are positioned so that the cells in the two layers are offset and interdigitate at their contiguous surfaces (see Noguchi, 1988: fig 115c). This feature is also seen in the leaves of Diphyscium Mohr and in the costa of Campylopus albidovirens Herz. Other distinctive features of the genus include its well-developed stem central strand, denticulate to serrate leaf margins that are incurved above when wet, and slender peristome teeth that are erect and sometimes twisted clockwise. Within the Pottiaceae, this last feature is present found only in Leptodontiella Zand. & E. Hegew. (Zander 1993).

Zander (1993) considered Timmiella a basal group within the Pottiaceae that shares many primitive characters with his out groups Polytrichum Hedw., Timmia Hedw, and Ptychomitrium Fürnr.


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Timmiella (De Not.) Limpr., Laub. Deutschl. 1: 590. 1888.

 Trichostomum sect. Timmiella De Not., Cronac. Briol. Ital. 1: 14. 1866. 

Plants dull, yellow-green to green, in dense or loose tufts. Stems erect, sparsely and irregularly branched, hyalodermis present, central strand well-developed. Leaves erect at base, incurved, spirally twisted and contorted, tubular above when dry, spreading when wet; apices acute; lamina bistratose, bistratose cells near costa at mid-leaf not vertically aligned; margins denticulate or serrate above, serrulate below, plane to incurved above; costa percurrent to short excurrent, wide, guide cells and two stereid bands well-developed, ventral surface layer of short, bulging-mammillose cells; upper cells rounded-quadrate, firm-walled, bulging-mammillose ventrally, plane dorsally, basal cells enlarged, thin-walled, bulging, hyaline, smooth, alar cells not differentiated. Dioicous or monoicous, perichaetial leaves weakly differentiated. Setae elongate, smooth, spirally twisted. Capsules narrowly cylindrical; opercula high-conic; annuli well- or poorly developed; peristome teeth divided nearly to the base into 32 slender, densely spiculose, filaments, straight or weakly twisted clockwise. Calyptrae cucullate.

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