Plants small to medium-sized, to 15 mm high, in dense tufts. Stems erect, usually branched, densely foliate. Leaves oblong-elliptic, often keeled, contorted with in-rolled margins when dry, erect-spreading when moist; margins plane, sometimes revolute in the lower half; costa stout, shortly excurrent as an apiculus, highly papillose on the back; upper leaf cells small, rounded-hexagonal, mammillose on both sides or papillose; basal cells larger, smooth, more or less hyaline. Gemmae global, consisting of 3–4 cells. Dioicous. Setae ca. 8 mm long; capsules erect, cylindrical, ca. 2.0–2.5 mm long.