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Published In: Bulletin of the Buffalo Society of Natural Sciences 32: 94. 1993. (Bull. Buffalo Soc. Nat. Sci.) Name publication detail

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Acceptance : Accepted
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Tuerckheimia valeriana has multicellular, dentate leaf margins and on the basis of this feature the species was originally placed in Leptodontium. Leptodontium differs from T. valeriana in having stems without a central strand and undifferentiated ventral surface cells on the costa. Ptychomitrium also has similar leaf margins, and Zander (1993) has suggested the species may belong in Ptychomitrium. Ptychomitium, however, does not have massive, multiplex leaf cell papillae. The species was transferred to Gymnostomum (Zander 1972) because of its stem central strand and differentiated ventral epidermal costal cells. It is presently placed in Tuerckheimia on the basis of its overall aspect, evenly thick-walled laminal cells, and massive papillae centered over the cell lumina.

Illustrations: Bartram (1929, Fig. 2 A–I); Bartram (1949, Fig. 47 J–M); Zander (1993, Pl. 14 16–17); Sharp et al. (1994, Fig. 188). Figure 89.
Habitat: Moist shaded bluffs by waterfall and on rotten logs; 900–2700 m.
Distribution in Central America: GUATEMALA. Quiché: Sharp 2399 (FH); San Marcos: Steyermark 35696a (F); COSTA RICA. San José: Croat 43440 (MO). PANAMA. Chiriquí: Crosby 3964 (MO, PMA).
World Range: Mexico; Central America.


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Tuerckheimia valeriana (Bartr.) Zand., Misc. Bryol. Lichenol. 8: 27. 1978.

Leptodontium valerianum Bartr., J. Wash. Acad. Sci. 19: 18. 1929. Gymnostomum valerianum (Bartr.) Zand., Bryologist 75: 277. 1972. Protologue: Costa Rica. Piedra Blanca, Province of San José, Costa Rica, altitude 2,400 meters, Oct. 9, 1927, Valerio 86

Plants small, caespitose or tufted, light-green to yellowish green, at times somewhat glaucous. Stems 2–3 cm high, sparsely and irregularly branched, central strand well-developed, sclerodermis present; rhizoids red-brown, smooth. Leaves evenly spaced, spreading when wet, spreading incurved from the base, weakly catenulate and twisted when dry, concave to keeled with a broad central groove, oblong to linear-lanceolate, 2–3 mm long; apices acute, apiculate; margins plane, coarsely and distantly dentate in upper half; costae percurrent to shortly excurrent, guide cells and two stereid bands present, ventral surface cells quadrate, papillose, dorsal surface cells small, thick-walled; upper cells homogeneous, subquadrate to hexagonal, 7–10 μm, at times short-rectangular near costa, walls firm to thick, pluripapillose, papillae massive, multiple, centered over the lumina, basal cells weakly differentiated, short-rectangular, 2–4:1 longer than wide, smooth, marginal cells not differentiated. Dioicous. Perichaetial leaves weakly sheathing otherwise not differentiated. Sporophytes unknown.



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