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Published In: Species Plantarum 2: 579. 1753. (1 May 1753) (Sp. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: I.C. Hedge
General/Distribution: About 30 species mainly in Europe and SW Asia. The prominently white-bearded thecae and the clearly trigonous nutlets are usually diagnostic for the genus.


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Annual or perennial herbs, herbaceous, with simple hairs. Leaves undivided, margins lobed, crenate or serrate, reniform to ovate, petiolate or sessite. Verticillasters many-flowered, in axils of upper leaves. Bracts small, linear. Flowers white, cream, pink or purplish. Calyx tubular or campanulate, not or scarcely bilabiate, 5-nerved; teeth ± equal, as long as or dearly longer than tube; tube internally glabrous or pilose. Corolla clearly longer than calyx, bilabiate; upper lip straight or hooded; lower lip equal to or shorter than upper, 3-lobed; tube exserted beyond calyx teeth, straight or curved, annulate or not within. Stamens 4, didynamous, anterior pair longer than posterior, included under upper lip of corolla; thecae bilocular, spreading, prominently bearded. Style clearly 2-lobed. Nutlets clearly trigonous below, obovoid, substipitate, laterally almost flanged, apically truncate-rounded, smooth, with or without irregular raised white blotches.

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1 Perennial with white flowers; upper stem leaves petiolate; corolla with a curved ± sturdy tube 1 Lamium album
+ Annual with pink-purple flowers; upper stem leaves amplexicaul; corolla with a slender tube 2 Lamium amplexicaule
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