Leafy erect perennial herbs. Stems simple or branched, glabrous or pilose. Leaves lobed or deeply laciniate, petiolate. Verticillasters many-flowered in upper leaf axils, remote or crowded into a spike-like inflorescence. Bracts subulate. Calyx turbinate, indistinctly bilabiate, 5-nerved; teeth 5, ± equal, spiny, spreading to reflexed. Corolla bilabiate, exserted; tube glabrous within or annulate; upper lip scarcely falcate; lower lip spreading or reflexed 3-lobed. Stamens 4, included, in two pairs; thecae 2-locular. Style equally bifid. Nutlets smooth, sharply triquetrous, clearly flat truncate at apex.