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Published In: Publications of the Field Museum of Natural History, Botanical Series 22(3): 216. 1940. (10 Sept 1940) (Publ. Field Mus. Nat. Hist., Bot. Ser.) Name publication detailView in Biodiversity Heritage Library
 

Project Name Data (Last Modified On 10/1/2020)
Acceptance : Accepted
Note : Tribe Sipaneeae
Project Data     (Last Modified On 10/2/2020)
Notes:

Steyermarkia includes one species found in a rather small area in in southern Mexico and northern Central America. The plants are hirsute, soft to suffrutescent herbs with the leaves clustered generally near the base. The stems are usually unbranched, but the plants produced adventitious roots and perhaps stoloniferous growth. This species has medium-sized oblanceolate leaves; persistent, ovate. interpetiolar, shortly bifid stipules; subcapitate to cymose inflorescences on elongated petioles; 4-merous flowers; well developed calyx lobes; showy, salverform, pink corollas; capsular fruits that open through the top portion; and small rounded seeds. The floral biology of this species is not known, and this species could be distylous. Only two flowering specimens have been seen, and both have anthers situated near the base of the corolla and the stigmas situated shortly below the corolla throat. The anthers were reported by Lorence et al. (2012) as dorsifixed, but they are basifixed on the material studied here. The capsules are distinctive in their dehiscence through a circular apical pore, which forms apparently by disintegration of the disk portion of the flower. Steyermarkia has been collected infrequently, in part because the region where it lives is not well explored but in part perhaps because it is not common. 

Steyermarkia has been included provisionally in Sipaneeae by Delprete based on morphological analysis, but it has not been studied with moledular data. This genus is disjunct geographically from the rest of the tribe, which is found in northern through easternl South America with one species ranging into Panama. Some other small Rubiaceae genera of enigmatic relationships are also found in this region, and the biogeographic history of this area is not yet well understood. Steyermarkia has not been studied in detail since its original description; for the most complete modern treatment see Lorence et al. (2012: 284), and a treatment and illustration were presented by Standley & Williams (1975). 

Steyermarkia is similar to some species of Amphidasya in growth form and general aspect; those Amphidasya species differ in their fimbriate or laciniate stipules, white corollas with the lobes valvate in bud, and baccate or at least indehiscent fruits.

Author: C.M. Taylor.
The content of this web page was last revised on 1 October 2020.
Taylor web page: http://www.mobot.org/MOBOT/Research/curators/taylor.shtml

Distribution: Evergreen forest at 0-600 m, at least often on limestone substrates, in southern Mexico and adjacent Guatemala.
References:

 

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Perennial herbs and subshrubs, unarmed, terrestrial, unbranched, with adventitious roots, without raphides in the tissues, densely hirsute to sericeous. Leaves opposite and often clustered near base, petiolate, entire, with higher-order venation not lineolate, without domatia; stipules interpetiolar and fused to petioles, ovate and shortly bifid, well developed, erect and perhaps imbricated in bud, persistent. Inflorescences axillary, subcapitate to shorty cymose, few- to several-flowered, pedunculate with the peduncles elongated and sometimes bearing foliaceous bracts along their length, bracteate. Flowers subsessile, bisexual, apparently homostylous, apparently protandrous, medium-sized and showy, perhaps fragrant and diurnal; hypanthium turbinate; calyx limb well developed, 4-lobed, without calycophylls; corolla salverform, pink, internally glabrous except sparsely to moderately puberulous-pilosulous with stout trichomes in throat and/or on bases of lobes, lobes 4, ovate and rounded, in bud imbricated, without appendage; stamens 4, inserted near base of corolla, anthers narrowly ellipsoid, basifixed, opening by linear slits, without appendage, included; ovary 2-locular, with ovules numerous in each locule, on axile placentas; stigmas 2, linear-oblong, included. Fruit capsular, subglobose to oblate, loculicidal through apical pore in disk area, leathery, somewhat small (ca. 0.8 cm diam.), smooth, with calyx limb persistent; seeds numerous, subglobose, small (0.3-0.4 mm long), foveolate.
 
 
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