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(Last Modified On 11/13/2012)
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(Last Modified On 11/13/2012)
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Genus
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ALCHEMILLA L.
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PlaceOfPublication
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Gen. PI. ed. 5, 58. 1754
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Reference
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L. M. Perry in Contr. Gray Herb. 84:1-57. 1929.
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Synonym
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Alchemilla, sect. Lachemilla Focke, in Engl. & Prantl, Nat. Pflanzenfam. 33:43. 1894. Lachemilla (Focke) Rydb. in N. Am. Fl. 22:380. 1908.
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Description
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Perennial herbs, often prostrate or procumbent, often stoloniferous. Leaves alternate, petiolate, usually palmately lobate or cleft, the upper smaller, with fewer lobes or divisions, sessile; stipules sheathing, usually foliaceous, lobed or cleft, adnate to the petiole, the whole often appearing as a multifid perfoliate leaf or a many- lobed sheath. Flowers minute, usually crowded into small axillary or terminal glomerulose cymes, less often in loose racemoid inflorescences; hypanthium per- sistent, urceolate, the 8-10 lobes 2-seriate, the outer ones smaller; petals none; stamens 1-4, free, opposite the calyx-lobes, the filaments short, usually not more than half the length of the calyx-lobes; disk with thickened margin, nearly closing the mouth of the hypanthium. Fruit of 1-6 achenes, these sessile or stipitate, in- cluded in the hypanthium, with basal, ventral, ascending and persistent styles, and capitate or clavate stigmas.
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Note
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This group of species, whether it be treated as a genus, Lachemilla, as done by Rydberg and more lately by Rothmaler (Fedde Rep. Sp. Nov. 42:164-173. 1937), or as a section or subgenus of Alchemilla, following Focke, Miss Perry, and more recent American authors including Standley (Field Mus. Publ. Bot. 18:477-478. 1937), and Standley & Steyermark (Fieldiana: Bot. 24:436-440. 1946), is easily recognized in the field, but the individual species are difficult of interpretation. The flowers are very small (3-4 mm. long or less), and in consequence the species have been distinguished chiefly by the use of characters of leaf-shape and lobing, distribution and quality of pubescence, and other vegetative characters which are not always convincing. Miss Perry divided the group as a whole into six easily recognizable series, or species-groups, of which two, the ORBICULATAE and the APHANOIDES, are represented in Panama.
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Key
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a. Basal leaves 5- to 11-lobed or -cleft, the blades relatively broad and merely toothed, even those of the upper leaves much broader than and distinct from the stipules. b. Leaf-blades shallowly 5- to 11-lobed- .......................... 1. A. ORBICULATA bb. Leaf-blades palmately 5- to 9-cleft to near their bases . -................... 2. A. PASCUORUM sa. Basal leaves 3- to 5-cleft or -parted, the blades with narrow finger-like lobes, those of the upper leaves often similar to and little exceeding the stipules-........... ...... . ... 3. A. APHANOIDES
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