Evergreen, small trees or shrubs, often spiny. Leaves simple, alternate, glandular punctate, petiole winged or margined. Flowers perfect or staminate, solitary or clustered in axillary racemes. Calyx 4-5-lobed, glabrous or pubescent. Petals (4-) 5(-8). Stamens 4-10 times the petals, polyadelphous. Ovary 10-14-locular, ovules biseriate or collateral. Fruit a fleshy hesperidium, globose to mamillate-oblong to oblate, rind tight or loose, with oil glands. Seeds embedded in pulpy vesicles.