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Published In: Species Plantarum 1: 543. 1753. (1 May 1753) (Sp. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: HARALD RIEDL and YASIN J. NASIR
Contributor Institution: Naturhistorisches Museum, Botanische Abteilung, Wien, Austria.


National Herbarium, Pakistan Agricultural Research Council, Islamabad, Pakistan.

General/Distribution: A genus of c. 250 cosmopolitan, but chiefly temperate species. Species with petaloid filaments or staminodes are sometimes separated in subgenus Atragene L.
Comment/Acknowledgements: Several species as Clematis hybrida Hort. and Clematis flammula L. are cultivated in gardens for their ornamental value. No poisonous plants are known in Clematis. Represented in Pakistan by 10-11 species.


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Climbing to erect shrubs, perennials rarely annuals. Leaves opposite, ternately or pinnately divided. Flowers solitary (-2-4) in fascicles or in paniculate cymes. Sepals 4-8, petaloid. Petals absent but filaments or staminodia sometimes petaloid. Stamens numerous. Carpels forming a small subglobose head. Ovules solitary, pendulous. Style often much elongating after flowering, plumose-hairy.

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1 Leaves pinnate, with very small lateral leaflets and much longer, simple, terminal leaflets
+ Leaves ternate to pinnate with ± equal leaflets (2)
2 (1) Leaflets linear-lanceolate to narrowly ovate, never cordate at base, sometimes distinctly lobed with entire lobes (3)
+ Leaflets with cordate base or ± broadly ovate, grossly dentate, rarely entire or lobed with a few grossly dentate lobes (6)
3 (2) Flowers single on long pedicels. Corolla-l ike sepals yellow, at least 3 cm long (4)
+ Flowers in panicles or few-flowered fascicles, not distinctly corolla-l ike, sepals white or pale yellow, less than 3 cm (5)
4 (3) Filaments dilated, petaloid, but always bearing fertile anthers 1 Clematis robertsiana
+ Petaloid staminodes present 10 Clematis alpina var. sibirica
5 (3) Filaments shorter than anthers 9 Clematis orientalis
+ Filaments longer than anthers 8 Clematis graveolens
6 (2) Leaves ternate. Flowers single or in fascicles on long pedicels (7)
+ Leaves pinnate or bi-pinnate (8)
7 (6) Sepals violet outside, narrower, acute to sub-acute. Anthers slightly bearded 11 Clematis barbellata
+ Sepals white to yellowish-white, obtuse to subacute, oblong to ovovate. Anthers glabrous 2 Clematis montana
8 (6) Leaflets neither leathery nor rigid (9)
+ Leaflets leathery, ± riged, often shiny on the upper surface. Flowers densely paniculate 3 Clematis gouriana
9 (8) Sepals 10-20 mm long-erect, curved backwards at the tips (10)
+ Sepals 6-8 (-10) mm long, horizontally spreading 4 Clematis grata
10 (9) Sepals with ± parallel ribs, 15-20 mm long 5 Clematis buchananiana
+ Sepals without ribs, 10-12 mm long 6 Clematis connata
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