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Project Name Data (Last Modified On 7/24/2013)

Flora Data (Last Modified On 7/24/2013)
Genus Themistoclesia Klotzsch
PlaceOfPublication Linnaea 24: 41. 1851.
Note LECTOTYPE: T. pendula Klotzsch.
Description Straggly to bushy, often epiphytic shrubs or shrublets. Leaves subcoriaceous to coriaceous, occasionally fleshy, alternate, shortly petiolate; venation obscure. Inflorescence axillary, racemose or solitary with one to several flowers, pedicels usually short, subtended by small bracts and bearing usually inconspicuous bracteoles. Flowers with the hypanthium continuous with the pedicel, obconic or short cylindric, narrowly 5-winged to the sinuses or bluntly angled, the calyx limb erect to slightly spreading, the 5 lobes acute to acuminate, minute to well developed; corolla ovoid to cylindric, the lobes 5; stamens 4-5 or 8-10, equal or alternately slightly unequal, more than 1/2 as long as the corolla and often about as long as the corolla tube, the filaments distinct, slender and terete to somewhat flattened at the base, the anthers inserted medially or almost basally, the walls smooth, nearly equal to or only about 1/2 as long as the tubules, the tubules 1 or 2, if 2, often completely separate, dehiscing by short, introrse clefts; style filiform, about as long as the corolla. Berry with numerous small seeds; the green embryo clearly visible through the surrounding tissue.
Habit shrubs or shrublets
Distribution Themistoclesia is a genus of about 25 species ranging from Costa Rica and Panama through the Andes from Venezuela and Colombia into Peru.
Key a. Staminal tubules 1 per anther with a single introrse cleft; hypanthium distinctly 5- winged; leaves apically obtuse ...... 3. T. pentandra aa. Staminal tubules 2 per anther, the tubules either separate or united but with 2 introrse clefts; hypanthium bluntly 5-angled, not winged; leaves apically acute or acuminate. b. Corolla 2.5-3.5 mm long; leaves less than 1.9 cm long, basally tapering or cuneate ...... 4. T. smithiana bb. Corolla 5-6 mm long; leaves more than 2 cm long, basally rounded, subtruncate or subcordate. c. Bracteoles ovate to hemispheric, completely concealing the glabrous pedi- cel; corolla cylindric, pilosulose throughout; stamens 5 mm long, filaments pilosulose, anther basally minutely appendiculate ...... 1. T. costaricensis cc. Bracteoles awl-shaped, never concealing the pilosulose pedicel; corolla urceolate, pilosulose distally; stamens 2.5 mm long, filaments glabrous, anther without basal appendage ...... 2. T. horquetensis
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