(Last Modified On 5/13/2013)
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(Last Modified On 5/13/2013)
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Genus
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Cornutia L.
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PlaceOfPublication
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Sp. P1. 628. 1753
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Synonym
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Agnanthus Vaill., Hist. Acad. Roy. Sci. [Gall.]. 1722: 201. 1724. Cornuta L., Gen. P1., ed. 1. 366. 1737; ed. 6. 316. 1764. Hosta Jacq., Hort. Schoenbr. 1: 60, pl. 114. 1797, non Hosta Tratt., Arch. Gewachsk. 2: 144. 1814, nom. cons. Hostana Pers., Syn. P1. 2: 143. 1806.
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Description
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Odoriferous trees and shrubs, mostly pubescent or tomentose throughout, often glandular-punctate; branches and branchlets mostly rather stout, tetragonal, often margined; nodes mostly annulate, often conspicuously so. Leaves simple, decus- sate-opposite, exstipulate, deciduous, petiolate, mostly large and broad, pubescent. Inflorescences mostly terminal, large pyramidal-thyrsoid panicles, sometimes also axillary and cymose, the cymes not involucrate. Flowers small, zygomorphic, perfect; calyx mostly small, cupuliform or cyathiform to campanulate, obconic, or patelliform, slightly irregular when mature, mostly unchanged or shrivelled to a patelliform condition in fruit, during anthesis entire or minutely 4(or 5)-dentate, rarely lobed; corolla hypocrateriform, zygomorphic, blue or violet to purple or white, the tube cylindric, often inflated at the base, constricted at the middle, ampliate above, straight or slightly curved, the adaxial side longer, pubescent within and often outside, the limb patent, 2-lipped, 4-parted, the adaxial lip 3-lobed with subequal ovate- or elliptic-lingulate lobes overlapping in bud or
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Habit
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trees and shrubs
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Description
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the central one outside, the abaxial lip composed of 1 lobe larger than the others and included by them in bud, made up of two distinct halves of which the upper is thinner, more glabrous, and folded in like a flap, the whole limb mostly pubescent inside and out; stamens 4, inserted in the corolla-tube at or above the middle, didynamous, the lower 2 fertile, longer, sometimes exserted, alternate with the largest corolla-lobe, the upper 2 reduced to filiform, mostly anantherous, included staminodes, the filaments flattened, mostly pubescent, the anthers 2- celled, dorsifixed, the thecae widely divergent basally, dehiscing by longitudinal slits, the connective much thickened; pistil 1, compound, mostly included, the style terminal, single, usually pubescent, of 2, 2-celled carpels, distinctly 4-celled, each cell with 1 high-lateral hemianatropous ovule; disk present beneath the ovary and about equalling it. Fruit drupaceous, small, abundant, subglobose, the exocarp fleshy, the endocarp hard and bony, the stones (1-3-)4-celled and -seeded; seeds without endosperm, the coat perforated at the base, stoney, rugose.
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Note
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A difficult genus of 25 mostly variable and polymorphic species and infra- specific entities of the American tropics. Many are handsome when in flower. The named species and varieties are in many cases not clearly defined.
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Key
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a. Branches, rachis, petioles and both leaf-surfaces short-pubescent, or pulverulent. b. Branches, branchlets, rachis, petioles, and lower leaf-surface densely short-pubescent. c. Flower-buds densely long hirsute -lb. C. grandifolia var. normalis cc. Flower-buds merely short-pubescent or strigose la. C. grandifolia var. intermedia bb.Branches, branchlets, and rachis scattered pilose, pulverulent or glabrate. d. Leaves villous with long, glistening, white, multicellular hairs ...... 2a. C. microcalycina var. anomala dd. Leaves short-pubescent or glabrate above, densely short-tomentose or pulverulent- puberulent beneath. e. Leaves densely or sparsely short-tomentose beneath - 2. C. microcalycina ee. Leaves merely pulverulent or pulverulent-puberulent beneath ...... 2b. C. microcalycina var. pulverulenta aa. Branches, rachis, petioles and both leaf-surfaces long-pubescent, hirsute, villous, or tomentose ...... 1. C. grandifolia
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