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Project Name Data (Last Modified On 6/6/2013)

Flora Data (Last Modified On 6/6/2013)
Genus Archibaccharis Heering
PlaceOfPublication Jahrb. Hamburg Wiss. Anst. Beib. 21(3): 40. 1904.
Note LECTOTYPE: A. hieraciifolia Heering.
Synonym Hemibaccharis Blake, Contr. U. S. Natl. Herb. 20: 552. 1924. T-YPE: H. hieracioides Blake.
Description Herbs or weak, often divaricate, scandent shrubs to 4(-10) m tall, mostly pubescent; stems often striate, sometimes fistulose. Leaves alternate, mem- branaceous or chartaceous, dentate in the upper portion or subentire with tooth- like, well spaced, callose mucros, pinnately veined, mostly pubescent, sometimes glandular, rarely scabrous; petioles mostly short or wanting. Inflorescence a panicle, often rounded, open or compact; reduced, leaflike bracts and scalelike bracteoles mostly present; pedicels filiform. Plants dioecious, perhaps sometimes polygamous; heads disciform, mostly with pistillate outer florets and hermaph- rodite inner florets but only one sex fertile on a plant; involucral bracts imbricate, in several unequal and unlike series, the innermost lanceolate, glabrous with a slender midvein, the outer series broader, often pubescent with a broader midvein, the hyaline margins apically erose and flanking an indurated stramineous region; receptacle flat, naked, smooth or alveolate; outer florets slender or capillary, the corolla cylindrical, hardly expanded at the base, apically truncate, erose, dentic- ulate or with a small ligule, often pubescent upwards, the style exserted, its branches slender, mostly elongate, glandular, the ovary compressed, pubescent, more so at the base, the pappus of fine strigose bristles in one series; inner florets with crateriform corollas, the corolla tube and base of the limb mostly pubescent, the limb tubular or campanulate, 5-lobed, sometimes deeply so, the lobes acute or ensiform with thickened margins, mostly glabrous, the anthers auricled on the inner (ventral) side, the appendages lanceolate, the style sometimes pubescent, its branches elongate, fusiform, glandular-papillose dorsally, mostly exserted from the anther appendages, the ovary sterile, mostly shrunken, the pappus slightly expanded apically. Achene compressed, often pubescent.
Habit Herbs shrubs
Note Archibaccharis is distinct in being dioecious while having sterile florets of the opposite sex in at least the pistillate heads. The leaves of most species are also distinctive in the minute teeth of callose mucros which stand at a prominent angle to the leaf margin. The genus differs from Baccharis in mostly lacking coriaceous leaves, in possessing two kinds of florets in the heads, in the presence of flat, smooth, or alveolate receptacles, and in the absence of a conspicuous swelling at the base of the corolla of the outer (pistillate) florets. Archibaccharis is also closely related to Conyza and Erigeron. Two interesting features possessed by the genus are the minute auricles on the inner (ventral) surfaces of the anther bases and the tendency of the staminate florets to change from white to rose or purplish in age, colors unusual in disc florets in this tribe.
Distribution This is a genus of about 20 species ranging from Mexico to Panama. The three Panamanian species are all from middle and upper elevations.
Reference Jackson, J. D. 1975. Revision of the genus Archibaccharis Heering (Compositae- Astereae). Phytologia 32: 81-194.
Key a. Styles of capillary outer florets exserted less than 1/3 their length; inner florets mostly more than 3.5 mm long; leaves mostly more than 3 times as long as broad. b. Capillary outer florets apically erose or truncate, without a defined ligule; veins 6 or more on each side of the midvein; limb of the inner crateriform florets divided less than halfway ...... 1. A. irazuensis bb. Capillary outer florets with a definite ligule ca. 1 mm long; veins ca. 5 on each side of the midvein; limb of the inner crateriform florets divided deeply ...... 2. A. panamensis aa. Styles of capillary outer florets exserted half their length or more; inner florets mostly less than 3.5 mm long; leaves mostly less than twice as long as broad ...... 3. A. schiedeana
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