Herbs or shrubs. Capitula homogamous discoid or heterogamous disciform or radiate. Involucral bracts often spinescent or foliaceous, many-seriate. Receptacle flat, naked. Ray florets when present, neuter, bilabiate or with a 3-lobed limb only. Disc florets with a narrow tube and wide limb. Anthers obtuse-acute or acuminate apically, sometimes with pilose tails. Style branches subapically pilose. Cypselas obovoid to obconical, pilose or glabrous. Pappus monomorphic, bristly or scaly or dimorphic with both bristles and scales.