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Cardamine leucantha O.E. Schulz

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Cardamine leucantha (Tausch) O. E. Schulz, Bot. Jahrb. Syst. 32: 403. 1903; Dentaria leucantha Tausch, Flora 19: 404. 1836. TYPE:

Cardamine cathayensis Migo, J. Shanghai Sci. Inst. Sect. III. 3: 223. 1937.

Cardamine koreana (Nakai) Nakai, Bull. Nat. Sci. Mus. Tokyo 31: 49. 1952. List bsionym.

Cardamine leucantha (Tausch) O. E. Schulz var. crenata D. C. Zhang, Bull. Bot. Res., Harbin 11: 42. 1991. TYPE: China, Anhui, Jingdexian, Yucun Wolonggou, 200 m, 7 May 1985, Shao Jianzhang 82143 (holotype, ANU).

Cardamine leucantha (Tausch) O. E. Schulz var. glaberrima F. Maekawa ex H. Hara, J. Jap. Bot. 51: 192. 1976.

Cardamine macrophylla Willdenow var. albiflora Makino, Bot. Mag. (Tokyo) 8: 302. 1894.

Cardamine macrophylla Willdenow var. parviflora Trautvetter & C. A. Meyer, Fl. Ochot. 15 1856.

Dentaria dasyloba Turczaninow, Bull. Soc. Imp. Naturalistes Moscou 27: 296. 1854; Cardamine dasyloba (Turczaninow) Miquel, Ann. Mus. Bot. Lugduno-Batavi 2: 73. 1866; D. macrophylla (Willdenow) Bunge ex Maximowicz var. dasyloba (Turczaninow) Makino, Bot. Mag. (Tokyo) 11: 157. 1897. TYPE:

Herbs, perennial, sparsely to densely villous. Rhizomes creeping, slender, not scaly or stoloniferous. Stems 2.5–7.5 dm, simple, flexuous. Cauline leaves 4–7, including petiole (8–)10–20(–25) cm; petiole (1–)2–8(–10) cm, not auriculate at base; terminal leaflet lanceolate, elliptic, to ovate-elliptic, (3–)4–9(–13) ´ (0.6–)1–3.5(–4) cm, adaxially shortly strigose or puberulent, abaxially pilose or long strigose, with a petiolule 5–13(–20) mm, base cuneate, margin antrorsely ciliate and irregularly serrate, double serrate, or subdentate, apex acuminate or rarely acute; lateral leaflets 2 or 3 pairs, rarely uppermost leaf trifoliolate, similar to terminal one, smaller, sessile, base cuneate or sometimes oblique. Racemes ebracteate, 12–24-flowered; fruiting pedicels divaricate or ascending, (0.5–)1–1.8(–2.3) cm, pilose or puberulent, slender. Sepals oblong, (2–)2.5–3.5 ´ 0.9–1.5 mm, margin membranous, pilose abaxially; petals white, spatulate to oblong-oblanceolate, 6–8 ´ 2–3.5(–4) mm, cuneate and not clawed at base, apex rounded; median filament pairs 5–6 mm, lateral pair 4–5 mm; anther oblong, 0.8–1.3 mm; ovules 6–12 per ovary. Fruit linear, (1–)1.5–3 cm ´ 1–1.5 mm; gynophore 0.3–0.8(–1) mm; valves smooth, sparsely hairy or glabrous; style slender, (2–)3–5 mm. Seeds brown, oblong, 1.5–2.2 ´ 0.8–1.2 mm, narrowly winged or wingless. 2n = 16.

Flowering: Apr–Jul.

Habitat: roadsides, shady areas, forests, wet places along streams, roadsides.

Elevation: 100–2000 m.

Distribution: China (Anhui, Gansu, Guizhou, Hebei, Heilongjiang, Henan, Hubei, Jiangxi, Jilin, Jiangsu, Liaoning, Nei Mongol, Ningxia, Shaanxi, Shanxi, Sichuan), Japan (Honshu, Hokkaido), Korea, Mongolia, Russia (Far East).