Cardamine douglassii Britton, Trans. New York Acad. Sci. 9: 8. 1889 Dentaria douglassii (Britton) Greene, Pittonia 3: 124. 1896. TYPE:

Arabis rhomboidea Persoon var. purpurea Torrey, Amer. J. Sci. 4: 66. 1822; Dracamine purpurea (Torrey) Nieuwland, Amer. Midl. Naturalist 4: 40. 1915. TYPE:

Cardamine douglassii var. albidula Farwell, Amer. Midl. Naturalist 9: 26. 1925. TYPE:

Thlaspi tuberosum Nuttall, xx. TYPE:

Herbs, perennial, hirsute throughout or glabrous proximally. Rhizomes fleshy, tuberous at stem base, subglobose, lobed or not, (3–)4–10 mm in diam.; stolons absent. Stems (0.7–)1–2.5(–3) dm, erect, simple, sparsely to densely hirsute with trichomes (0.2–)0.3–0.6(–0.8) mm, or glabrous near base. Rhizomal leaves simple, (3–)5–15(–18) cm; petiole (2–)4–12(–16) cm; blade orbicular to cordate, sometimes reniform or ovate, (1–)2–6 × (0.7–)1.7–5 cm, base obtuse to cordate, margin repand or entire; cauline leaves 3–6(–8), simple, not auriculate at base, middle ones short petiolate, 2–5 × 0.5–2.5 cm, upper leaves sessile, oblong to ovate or lanceolate, entire, repand, or coarsely dentate. Racemes ebracteate; fruiting pedicels ascending to divaricate, (1–)1.5–3.5(–5) cm, sparsely pubescent or glabrous. Sepals oblong, 2.5–4(–6) × 1.5–2.5 mm, erect, often hirsute, base of lateral pair not saccate; petals rose-purple to pink of very rarely white, obovate, (7–)8–13(–15) × 3–5 mm, short clawed, apex rounded; median filament pairs 4–7 mm, lateral pair 2–4 mm; anthers oblong, 1.3–1.7 mm. Fruit linear, (1.5–)2–4 cm × 1.5–2 mm; valves glabrous; style 2–5 mm; ovules and seeds 10–16 per fruit. Seeds brown, oblong to ovate, 1.7–2.5 mm × 1–5 mm. 2n = 56, 64, 96, 112, 144.

Flowering: Mar–May.

Habitat: rich woods, bluffs, mesic bottomland foress, rocky hillsides, floodplains, seepage of bogs, springy areas.

Elevation: 50–400 m.

Distribution: Canada (S Ontario), United States (Alabama, Connecticut, District of Columbia, Illinois, Indiana, Iowa, Kentucky, Maryland, Massachusetts, Michigan, Minnesota, Missouri, New Jersey, New York, North Carolina, Ohio, Pennsylvania, South Carolina, Tennessee, Virginia, West Virgina, Wisconsin).