Cardamine clematitis Shuttleworth ex A. Gray, Proc. Amer. Acad. Arts 15: 45. 1880. TYPE:

Herbs, perennial, glabrous throughout. Rhizomes slender, cylindrical, 1–3 mm in diam.; stolons absent. Stems (0.8–)1–2.5(–3.5) dm, erect, simple or rarely few branched above, glabrous. Rhizomal leaves simple or 3-foliolate, (1.5–)3–8 cm; petiole (1–)2–6 cm; blade or terminal leaflet reniform to cordate, (0.5–)1–2 cm, petiolule to 1 cm, base cordate, margin entire or 3-lobed, not puberulent; lateral leaflets much smaller than terminal one, subsessile, margin same as terminal leaflet; cauline leaves 3–7, 3-foliotate or uppermost simple; petiole 0.7–3.5 cm, base auriculate, the auricle 0.7–5 mm; terminal lobe broadly ovate to suborbicular or reniforem, 1.5–4 × 1.5–3.5 cm, often 3–5-lobed, the lobe apex minutely apiculate, glabrous along margin, petiolule 0.5–1 cm; lateral leaflets oblong to ovate or oblong, sessile or on petiolule to 0.5 cm. Racemes ebracteate; fruiting pedicels divaricate-ascending, (0.7–)1–1.7 cm. Sepals oblong, 2.5–3 × 1–1.5 mm, erect, base of lateral pair not saccate; petals white, oblanceolate, 6–8 × 2–3 mm, not clawed, apex obtuse to subemarginate; median filament pairs 3.5–4 mm, lateral pair 2–2.5 mm; anthers ovate, ca. 0.7 mm. Fruit linear, (1.5–)2–3.5–(4) cm × 1.3–1.7 mm; valves glabrous; style 2–4 mm; ovules and seeds 10–16 per fruit. Seeds brown, oblong, 1.7–2 × 1–1.2 mm.

Flowering: May–Jun.

Habitat: wet areas, springs, moist slopes.

Elevation: 1300–1800 m.

Distribution: United States (Alabama, Florida, Georgia, North Carolina, Tennessee, SW Virginia).