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Published In: Genera Plantarum 90. 1789. (4 Nov 1789) (Gen. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: YASIN J. NASIR
Contributor Institution: National Herbarium, Agricultural Research Council, Islamabad.
General/Distribution: A family of 30 genera and c. 290 species, in tropical and subtropical parts of both the hemispheres, especially America. Represented in Pakistan by 5 genera and 10 or 11 species. In the key that follows and the descriptions, the term anthocarp stands for the mature fruit and ‘lower half of perianth’ for the anthocarp at anthesis.
Comment/Acknowledgements: Acknowledgements: We are grateful to the United States Department of Agriculture for financing this research under P.L. 480. Thanks are also due to Dr. F.R. Fosberg, Smithsonian Institution, Washington, Mr. R.D. Meikle, Herbarium, Royal Botanic Gardens, Kew, Mr. B.L. Burtt, Mr. I.C. Hedge and Miss J. Lamond of the Royal Botanic Garden, Edinburgh for going through the manuscript and giving valuable suggestions.


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Trees, herbs or shrubs. Leaves opposite or alternate, simple, exstipulate. Inflorescence cymose. Flowers hermaphrodite or unisexual, often bracteate; bracts showy and petaloid or inconspicuous. Perianth 5-lobed, tubular, gamophyllous, petaloid, lower half (the anthocarp) persistent. Stamens 5, but may be, more or less, alternating with the lobes; filaments often unequal, anthers bilocular, dehiscing longitudinally. Pistil monocarpellary, ovary unilocular, with a solitary basal ovule. Fruit a nut, utricle or an achene, enclosed in the anthocarp. Seed endospermic.

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1 Annual or perennial, prostrate or erect diffuse herbs, rarely shrubs, up to 1.5 m tall (2)
+ Trees or shrubs, climbing or often straggling, up to 12 m or more tall (4)
2 (1) Flowers small, bracts or involucel of bracts not prominent (3)
+ Flowers showy, with an involucre of prominent bracts Mirabilis
3 (2) Anthocarp 10-ribbed, with large wart-like glands Commicarpus
+ Anthocarp 5-ribbed, glabrous or glandular, without wart-like glands Boerhavia
4 (1) Usually medium sized trees. Flowers without conspicuous bracts. Anthocarp viscous, glandular Pisonia
+ Straggling or climbing shrubs. Flowers with 3 conspicuously coloured bracts. Anthocarp glabrous or pubescent, never viscous Bougainvillea
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