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Published In: Genera Plantarum 163. 1789. (4 Aug 1789) (Gen. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: E. NASIR
Contributor Institution: National Herbarium, Pakistan Agricultural Research Council, Islamabad.
General/Distribution: A large family with about 40 genera and 600 species, mainly in the northern hemisphere. Represented in Pakistan by 3 genera and c.26 native species.
Comment/Acknowledgements: Acknowledgements: We are grateful to the U.S. Department of Agriculture for financing this research under PL-480. We are also thankful to the Directors of the Herbaria, Royal Botanic Gardens at Kew and at Edinburgh for the loan of herbarium specimens and ,to Mr. I.C. Hedge for going through the manuscript and the latin diagnosis of the new species of Campanula.


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Annual or perennial herbs, sometimes lactiferous. Roots and rhizomes often thickened and tuberous. Leaves simple, usually alternate, sometimes opposite, exstipulate. Stems erect, decumbent, trailing or climbing. Flowers usually hermaphrodite, actinomorphic, regular, sometimes cleistogamous. Calyx-tube adnate to the ovary. Sepals with linear to triangular or lanceolate segments. Corolla united, campanulate, infundibuliform or tubular, 5-lobed, sometimes the segments split to the base. Stamens 5 with flattened filaments. Ovary inferior, (2-)3-5 locular, with numerous ovules; styles thick with 3, sometimes 2 stigmas. Fruit a capsule, dehiscing by valves or pores.

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1 Fruit capsular, not beaked, dehiscence porous by rupture of the fruit wall at the apex, middle or the base. Plants erect, decumbent or trailing. (2)
+ Fruit a beaked capsule, dehiscence by 3 loculicidal valves at the apex. Plants twiners, decumbent or erect. 1 Codonopsis
2 (1) Plants erect. decumbent or trailing. Corolla campanulate, funnel-shaped, tubular, lobes short, less than half the length of the corolla. 3 Campanula
+ Plants erect. Corolla not campanulate, lobes split to the base of the corolla into narrow segments. 2 Asyneuma
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