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  • Reference article Robbrecht, E. & J. F. Manen. 2006. The major evolutionary lineages of the coffee family (Rubiaceae, angiosperms). Combined analysis (nDNA and cpDNA) to inferthe position of Coptosapelta and Luculia, and supertree construction based on rbcL, rps16, trnL-trnF and atpB-rbcL data. A new classification in two subfamilies, Cinchonoideae and Rubioideae. Syst. & Geogr. Pl. 76: 85–146.
  • Reference article Taylor, C. M. 2010-2025. Studies in Tropical Rubiaceae.
  • Reference article Torres-Montúfar, A., T. Borsch, S. Fuentes-Bazán, J. Gutiérrez & H. Ochoterena-Booth. 2020. It is not a disaster: molecular and morphologically based phylogenetic analysis of Rondeletieae and the Rondeletia complex (Cinchonoideae, Rubiaceae). Pl. Syst. Evol. 306(26): 1–25.
 
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