Accepted By:
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Alejandro, G. D. & S. Liede-Schumann. 2003. The Philippine Rubiaceae Genera: Updated Synopsis in INTKEY Databases of the DELTA System. Blumea 48: 261-277.
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Biag, R. D. & G. D. Alejandro. 2021. Rubiaceae flora of nothern Sierra Madre National Park, Isabela, Luzon, Philippines: Species richness, distribution, and conservation status. Philipp. J. Sci. 150(3): 907–921.
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