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Published In: Enumeratio Plantarum Omnium Hucusque Cognitarum 2: 394. 1837. (1-6 May 1837) (Enum. Pl.) Name publication detailView in Biodiversity Heritage Library
 

Project Name Data (Last Modified On 9/1/2009)
Acceptance : Accepted
 

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18. Section Ovales (Kunth) H. Christ

Plants monoecious, mostly with short-creeping rhizomes, forming loose to dense tufts or clumps. Vegetative stems present, variously short or elongate, sometimes reduced to basal clusters of leaves or nearly so. Flowering stems erect, mostly longer than the leaves, sharply to bluntly trigonous, unwinged, firm, glabrous, smooth or roughened on the angles near the tip, green or light brown to dark brown at the base. Leaves basal and on the basal half of the stems, glabrous, the basal leaves reduced to nearly bladeless sheaths, the brown remains of the lowermost leaves persisting through the following year. Leaf blades flat or somewhat folded near the base, the margins minutely roughened or toothed. Leaf sheaths green or sometimes pale green to white with green lines near the tip, the ventral side firm or papery, green, white, or yellowish white, not cross-wrinkled or red-spotted, the lowermost sheath bases green or light brown to dark brown. Inflorescences with 2–15 spikes, these sessile or very short-stalked, widely or closely spaced on the axis, the bracts inconspicuous, scalelike, the lowermost sometimes with an elongate awn, shorter than the spikes, lacking a sheath. Spikes with a broad pistillate region toward the tip and a much narrower staminate region toward the base, this inconspicuous or prominent, occasionally the lowermost spikes entirely pistillate, elliptic to obovate or nearly circular in outline, the base and tip appearing rounded or pointed, with numerous densely spaced perigynia, these ascending or spreading at maturity, sometimes radiating in all directions. Staminate and pistillate scales similar, the staminate scales usually slightly longer than the pistillate ones. Perigynia thin-walled, flat to somewhat concave ventrally, flat to somewhat rounded dorsally, ovate to narrowly elliptic-lanceolate in outline, tapered to a flattened beak with minute teeth along the margins and 2 narrow teeth at the tip, the base rounded or tapered, the sides mostly strongly winged, glabrous. Styles withering during fruit development, jointed to the main body of the fruit, which is not or minutely beaked at maturity. Stigmas 2. Fruits biconvex and somewhat flattened in cross-section, shiny. About 85 species, North America to South America, Hawaii, Europe, Asia; introduced in New Zealand.

Section Ovales is among the most easily recognized groups of Missouri sedges, but the species within the section can be very difficult to determine. The section is one of the largest and most difficult taxonomically in North America. To help with determinations of specimens of Ovales, collectors should note characteristics of the vegetative stems, which can be important in distinguishing closely related species. When examining spikes, it is important to locate the most mature, best developed perigynia in a sample, as these facilitate interpretation of subtle characters of perigynium shape, size, and texture.

 

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1 Perigynia 2 mm wide or less, usually (but not always) less than 4 mm long (2)
+ Perigynia more than 2 mm wide, often (but not always) more than 4 mm long (13)
2 (1) Perigynia (including beaks) 2.5–5.0 times as long as wide, narrowly to broadly lanceolate (3)
+ Perigynia (including beaks) about as long as wide or up to 2.5 times as long as wide, broadly lanceolate to broadly elliptic-ovate in outline (7)
3 (2) Vegetative stems usually few at flowering time, mostly developing late in the growing season, the leaves clustered toward the tip; leaf blades 1–4 mm wide; perigynia winged to the base, the wings sometimes gradually narrowed toward the base; distance from the tip of the fruit to the tip of the perigynium 2.2–5.0 mm (4)
+ Vegetative stems abundant at flowering time, the leaves well spaced along the apical half; leaf blades 3–8 mm wide; perigynia with the wings usually abruptly narrowed or absent toward the base; distance from the tip of the fruit to the tip of the perigynium 0.8–2.2 mm (6)
4 (3) Perigynia 0.8–1.2 mm wide, 3.5–5.0 times as long as wide 62 Carex crawfordii
+ Perigynia 1.3–2.0 mm wide, 2.5–3.5 times as long as wide (5)
5 (4) Fruits 0.7–0.8 mm wide; spikes mostly pointed at the tip, ovate-elliptic in outline, gradually tapered to the staminate base 70 Carex scoparia
+ Fruits 0.9–1.3 mm wide; spikes mostly rounded at the tip, obovate in outline, abruptly narrowed to the staminate base 73 Carex tenera
6 (3) Perigynia with the tips spreading stiffly in all directions, the spike appearing as a spiny ball; pistillate scales mostly hidden by the perigynia 63 Carex cristatella
+ Perigynia ascending or less commonly loosely spreading, but not stiffly spreading in all directions; pistillate scales noticeable and not hidden by the perigynia 74 Carex tribuloides
7 (2) Pistillate scales tapered to a conspicuous, slender, sharp point, usually awned 72 Carex straminea
+ Pistillate scales rounded to sharply pointed, but not tapered to a slender point or awned (8)
8 (7) Main body of the perigynium broadly obovate to nearly circular in outline, broadest above the middle 59 Carex albolutescens
+ Main body of the perigynium narrowly lanceolate to broadly ovate or more or less circular in outline, broadest at or below the middle (9)
9 (8) Main body of the perigynium more or less circular in outline, rounded abruptly to a narrow beak 64 Carex festucacea
+ Main body of the perigynium lanceolate to broadly ovate-elliptic, mostly tapered into a broadly based beak; species often difficult to distinguish (10)
10 (9) Inflorescences mostly stiffly erect; spikes mostly overlapping along the axis (internode between lowermost spikes mostly 2.5–6.0 mm); leaves with well-developed blades mostly 3–7 per flowering stem, the leaf blades 2–6 mm wide (11)
+ Inflorescences mostly curved or arched from just above the lowermost spike; spikes mostly loosely spaced along the axis (internode between lowermost spikes mostly 6–21 mm); leaves with well-developed blades mostly 3 or 4 per flowering stem, the leaf blades 1.5–3.5 mm wide (12)
11 (10) Perigynia pale brown to light tan at maturity, the main body broadly ovate-elliptic in outline; leaf blades 2–3 mm wide 65 Carex molesta
+ Perigynia usually green at maturity, sometimes turning straw-colored with age, the main body lanceolate to narrowly ovate in outline; leaf blades 2.5–6.0 mm wide 68 Carex normalis
12 (10) Internode between lowermost spikes mostly 6–11 mm, the axis relatively stout and stiff; perigynia mostly 1.8–2.2 times as long as wide 68 Carex normalis
+ Internode between lowermost spikes (6–)10–21 mm, the axis relatively slender and wiry; perigynia mostly 2.1–2.8 times as long as wide 73 Carex tenera
13 (1) Spikes 15–25 mm long, narrowly elliptic-lanceolate in outline, sharply pointed at the tip; perigynia 6.5–10.0 mm long, narrowly lanceolate in outline 66 Carex muskingumensis
+ Spikes 5–16 mm long, obovate to nearly circular or elliptic-ovate in outline, rounded to bluntly pointed at the tip and/or the base (mostly pointed in C. scoparia); perigynia 2.4–8.0 mm long, if longer than 5.5 mm then of a broader shape than lanceolate (14)
14 (13) Main body of the perigynium broadly obovate to nearly circular in outline, broadest above the middle (15)
+ Main body of the perigynium narrowly lanceolate to broadly ovate or more or less circular in outline, broadest at or below the middle (16)
15 (14) Pistillate scales rounded to sharply pointed, but not tapered to a slender point or awned 59 Carex albolutescens
+ Pistillate scales tapered to a conspicuous, slender, sharp point, usually awned 58 Carex alata
16 (14) Perigynia with the tips ascending, mostly appressed (17)
+ Perigynia with the tips loosely ascending to spreading, not appressed (20)
17 (16) Spikes rounded at the tip, circular to broadly ovate or obovate in outline (18)
+ Spikes mostly pointed at the tip, elliptic to elliptic-ovate in outline (19)
18 (17) Perigynia nerveless or with faint, short nerves near the base of the ventral side 61 Carex brevior
+ Perigynia with 6–9 distinct nerves on the ventral side 66 Carex molestiformis
19 (17) Ventral side of the leaf sheaths with an elongate, thinner, white area; spikes tapered gradually to the staminate base 70 Carex scoparia
+ Ventral side of the leaf sheaths green nearly to the tip, the thinner, white area short; spikes narrowed abruptly to the staminate base 72 Carex suberecta
20 (16) Largest perigynia 5.5–8.0 mm long, the beak 1.5–3.0 mm long (21)
+ Largest perigynia 2.5–5.5 mm long, the beak 0.8–1.5 mm long (2.0–2.5 mm long in C. straminea) (22)
21 (20) Perigynia with the main body wider than long, the surface finely granular with minute papillae (visible with magnification) 69 Carex reniformis
+ Perigynia with the main body as long as wide or slightly longer than wide, the surface smooth 60 Carex bicknellii
22 (20) Main body of the perigynium lanceolate to narrowly ovate in outline; leaves 2.5–6.5 mm wide 68 Carex normalis
+ Main body of the perigynium broadly ovate-elliptic to broadly elliptic or nearly circular in outline; leaves 1.5–4.0 mm wide (23)
23 (22) Perigynia with 3–7 definite nerves on the ventral surface, at least some of these as long as the fruit or longer (24)
+ Perigynia nerveless on the ventral surface, or if faintly nerved then the nerves shorter than the fruit (27)
24 (23) Pistillate scales tapered to a conspicuous, slender, sharp point, usually awned 71 Carex straminea
+ Pistillate scales rounded to sharply pointed, but not tapered to a slender point or awned (25)
25 (24) Main body of the perigynium more or less circular in outline; terminal spike usually narrowed to a prominent, staminate base; inflorescences mostly with 5–9 spikes 64 Carex festucacea
+ Main body of the perigynium broadly ovate-elliptic in outline; terminal spike usually with an inconspicuous, staminate base; inflorescences mostly with 3–5 spikes (26)
26 (25) Fruits 0.9–1.3 mm wide, 1.3–1.6 times as long as wide, elliptic to narrowly oblong in outline 65 Carex molesta
+ Fruits 1.4–1.8 mm wide, 1.0–1.3 times as long as wide, broadly oblong to nearly circular in outline 66 Carex molestiformis
27 (23) Perigynia with the main body 3.2–5.0 mm wide, wider than long, the surface finely granular with minute papillae (visible with magnification) 69 Carex reniformis
+ Perigynia with the main body 1.5–3.5 mm wide, as long as wide or slightly longer than wide, the surface smooth (28)
28 (27) Fruits 1.7–2.0 mm long; perigynia 2.3–3.5 mm wide 61 Carex brevior
+ Fruits 1.3–1.7 mm long; perigynia 1.5–2.4 mm wide 64 Carex festucacea
 
 
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