Shrubs and small trees, unarmed, terrestrial, without raphides in the tissues, resinous at apices. Leaves opposite, petiolate, entire, with tertiary and quaternary venation not lineolate and usually not evident, with pubescent domatia; stipules interpetiolar, elliptic to ovate or obovate, generally held erect and flatly pressed together in bud, quickly deciduous. Inflorescences axillary at 1--2 upper stem nodes, cymose, multiflowered, pedunculate, bracts reduced or developed and deciduous. Flowers subsessile or pedicellate, bisexual, distylous, protandrous, medium-sized, fragrant, diurnal or perhaps noctural; hypanthium ellipsoid; calyx limb developed, 5-lobed, without calycophylls; corolla salverform, white, internally pubescent, lobes 5, triangular, valvate in bud, with terminal appendages; stamens 5, inserted in corolla tube, anthers narrowly oblong, dorsifixed near base, included or partially exserted, opening by longitudinal slits, without appendages; ovary 2-locular, ovules numerous in each locule, imbricated and ascending on axile placentas, stigmas 2, linear, included or perhaps sometimes partially exserted. Fruit capsular, ellipsoid, septicidally dehiscent from apex, chartaceous, smooth, with calyx limb persistent or tardily deciduous; seeds numerous, flattened, small, fusiform to elliptic, marginally winged and fimbriate to ciliate.