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Published In: Bulletin of Miscellaneous Information, Royal Gardens, Kew 1894: 340. 1894. (Bull. Misc. Inform. Kew) Name publication detailView in Biodiversity Heritage Library
 

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Flower/Fruit: Fl. Per.: October-November.
Type: Lectotype: [S Yemen] Shari Burrock Valley, Mokalla, 200-300’, 21.12.1893, Lunt 53; (K!), iso- (BM!, P!). Botschantzev, l.c. 1975.
Habitat: Common and forming communities of its own on gypsiferous sands, pebbles, weak sedimentary rocks, in washes, on higher playa edges and on eroded slopes in the most severe hot deserts.By its smooth hairs and very remarkable disc the species stands rather apart from sympatric S. imbricata, but in habit sometimes it looks rather similar. Can easily be identified by its rigid branches, usually pungent; appearance whitish from the dense indumentum on all leafy organs; without fishy smell. The species is very palatable, but due to the intricate spiny branch system it is browsed almost exclusively by camels.
Distribution: Distribution: From Egypt through the Arabian Peninsula and S Iran to Pakistan.

E Saharo-Sindian.

Map Location: F-2 Makran: 47 km NW Panjgur, Freitag & Kothe 18682 (KAS); 60 km SSW Panjgur, Freitag & Kothe 18596 (KAS); 20 km from Parom to Panjgur, Ghafoor & Omer 1925 (KUH); 140 km from Buleda to Panjgur, Ghafoor & Omer 1876 (KUH); F-2/3 Makran: New road to Hoshab, 20 km WSW Panjgur, Freitag & Kothe 18623 (KAS); Ghafoor & Omer 1896 (KUH); 93 km S Panjgur, Freitag & Kothe 18588 (KAS); F-3 Kharan: 10 km W Palantak, road to Qila Ladgasht, Freitag & Kothe 18685a (KAS); Makran: near Panjgur, 900 m, Popov 146 (BM); 80 km from Awaran on road to Hoshab, Malik & Nazimuddin 1101 (KUH); Bela to Awaran, 35 km W Lak pass, Freitag & Kothe 18269 (KAS); 27 km W Lak pass, Freitag & Kothe 18259 (KAS); Ghafoor & Omer 1703 (KUH); 30-50 km W Bela, 200 m, Rechinger 27624 (W); Lamond 318 (E, G-PAE); F-4 Kalat: 12 km S Khuzdar to Bela, Freitag & Kothe 18937 (KAS); Ghafoor & Omer 2235 (KUH); G-1 Makran: 48 km N Suntsar to Mand, Freitag & Kothe 18568 (KAS); G-2 Makran: 8 km S Hoshab to Pasni, Freitag & Kothe 18344 (KAS); Ghafoor & Omer 1768A (KUH); 28 km SE Turbat to Pasni, Freitag & Kothe (18392) (KAS); Makran: 6 km N Pasni to Turbat, Freitag & Kothe 18493 (KAS); 6 km WNW Pasni, Freitag & Kothe 18420 (KAS); c.7 km from Pasni to Gwadar, Ghafoor & Omer 1787 (KUH); 80-100 km S Turbat to Pasni, Rechinger 27850 (W); G-2/3 Las Bela: 95 km E Pasni, near Makola, Freitag & Kothe 18435 (KAS); Ghafoor & Omer 1838A (KUH); G-3 Las Bela: 117 km E Pasni, 20 km E Makola, Freitag & Kothe 18436 (KAS); Ghafoor & Omer 1754 (KUH)(a glabrous form); 43 km W Ormara, Freitag & Kothe 18443 (KAS); 20 km NW Ormara, Freitag & Kothe 18477, 18478 (KAS).

 

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Subshrub or small shrub, 20-60 cm high; white with spreading (axes) and adpressed (outer side of leaves, bracts, bracteoles) 0.3-0.6 mm long smooth, thick-walled, flat hairs, rarely sparsely hairy to almost glabrous. Stem intricately branched from the to 2cm thick base, gnarled and light grey like the branches; branches ascending to spreading, stiff, around the leaf bases with rings of persisting long hairs, becoming + spinescent, developing from gemma-like buds, cream-coloured; last order branches forming condensed, often regularly arranged 5-15 x 2.5-3.5(4) mm spikes. Leaves scale-like, broadly triangular, obtuse, 1.0-1.6 mm long and wide, at base dorsally much thickened, upper surface concave, glabrous; leaf axils on long shoots with silvery, angled, globular gemmae. Bracts and bracteoles subequal, similar to the leaves but only c. 1 mm long and wider than long. Tepals ovate, 1.4-1.6 mm long, the outer 1-1.2 mm wide, 1-veined, transverse line at 1/3, green blotch covered by the densely adpressed indumentum. Anthers 0.7-0.9 mm long including the 0.15-0.2 mm long obtuse appendage, divided for c. 2/3; filaments linear, 1.2-1.5 mm long, inserted on outer base of disc; disc cup-like, thickened, 0.3 mm wide, glabrous, the margin undulate or lobed. Style 0.4-0.6 mm long; stigmas 0.4-0.6 mm long, flat, inside shortly papillose. Fruiting perianth 4-5.5 mm diam., the wings subequal, pale, fresh often pinkish; upper parts of tepals stiff, straight, forming a cone; lower part of tepals moderately hardened, forming a broad obconical structure, flat base very narrow. Seed semi-globular, flattened above, 1 mm diam.
 
 
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