2. Hieracium gronovii L. (beaked hawkweed)
H. gronovii var. foliosum Michx.
H. floridanum Britton
Pl. 256 a, b;
Map 1068
Plants with a
short, usually erect or ascending rootstock, rarely with a short, spreading
rhizome. Stems mostly solitary, 30–85(–120) cm long, moderately to densely
pubescent, at least toward the base, with light orangish brown, spreading to
loosely ascending hairs 4–9 mm long having a bulbous or slightly expanded base,
these sometimes becoming sparse or nearly absent toward the tip, usually also
inconspicuously pubescent with cobwebby, minute, branched hairs, often also
with sparse, gland-tipped hairs toward the tip. Basal leaves usually persistent
at flowering, sessile to short-petiolate, the blade 3–20 cm long, narrowly
oblanceolate to obovate, rounded to sharply pointed at the tip, the surfaces
and margins pubescent with sparse to moderate, spreading, bulbous-based hairs
and often also sparse to moderate, minute, inconspicuous, branched hairs. Stem
leaves sometimes few and nearly basal, sometimes several and well spaced,
similar to the basal leaves but gradually reduced in size, more often sessile,
oblong-obovate to oblanceolate, the uppermost with the base often slightly
clasping the stem. Inflorescences elongate cylindrical panicles or racemes.
Involucre 6–9 mm long, the inner series of bracts narrowly oblong-lanceolate,
pubescent with inconspicuous, cobwebby, branched hairs and sometimes also
longer, spreading, usually dark-colored, gland-tipped hairs, the outer series
much less than 1/2 as long as the inner series. Ligulate florets 20–40.
Corollas 7–9 mm long, light yellow to yellow. Pappus bristles 4–5 mm long,
light yellowish to orangish brown. Fruits 2.5–4.0 mm long, somewhat tapered to
the slightly expanded tip. 2n=18. May–October.
Scattered,
mostly south of the Missouri River, absent from all but the eastern portion of
the Glaciated Plains Division (eastern U.S. west to Illinois, Kansas, and
Texas; Canada, Mexico, Central America, Caribbean Islands). Mesic to dry upland
forests, ledges and tops of bluffs, and borders of glades; also pastures, old
fields, and roadsides.
There has been
some controversy about the correct application of the name H. gronovii,
but the epithet was conserved at an international botanical congress with a
type specimen that fixes the application to the traditionally accepted
widespread species (Greuter et al., 2000). Specimens with more stem leaves and
relatively few-flowered inflorescences occasionally have been misdetermined as H.
scabrum.
Steyermark
(1963) corrected earlier reports for Missouri of the eastern H. venosum L.,
annotating the several specimens in question to H. gronovii based on
their apically somewhat tapered (vs. cylindrical) fruits. This was confirmed
during the present study. Hieracium venosum should continue to be
excluded from the Missouri flora. Hieracium gronovii is relatively
variable in plant height, as well as number and size of the stem leaves. One of
these specimens, a historical collection from Newton County, has unusually
narrow leaves and relatively long, spreading hairs. Possibly, it represents a
hybrid with H. longipilum. In general, the two species do not grow
together, but they are known to hybridize regularly in other parts of their
ranges.