8. Bidens laevis (L.) Britton, Sterns & Poggenb. (showy bur marigold)
Pl. 273 c, d;
Map 1156
Plants annual
(sometimes perennial farther south), terrestrial or occasionally emergent
aquatics, sometimes with taproots. Stems 20–80(–150) cm, erect to loosely
ascending, sometimes from a spreading base, occasionally rooting at the lower
nodes, glabrous. Leaves all more or less similar, sessile, opposite or rarely
the lowermost leaves in whorls of 3, the blade 2–15 cm long, oblong-lanceolate
to oblanceolate in outline, less commonly obovate or linear, unlobed, narrowed
or tapered at the base, tapered to a sharply pointed tip, the margins usually
sharply and finely to coarsely toothed, rarely nearly entire, also minutely
roughened, the surfaces glabrous. Inflorescences of solitary terminal heads or
appearing in loose, open clusters, the heads radiate (rarely discoid
elsewhere), usually nodding at fruiting. Involucre with the outer series of 5–9
bracts 6–16 mm long, spreading to more commonly reflexed, somewhat leaflike,
narrowly lanceolate to narrowly oblong or linear, the margins entire but
usually minutely roughened, the outer surface glabrous or sparsely short-hairy
toward the base; the inner series of 6–9 bracts 4–10 mm long, lanceolate to
narrowly ovate or ovate, glabrous. Chaffy bracts narrowly oblong, reddish brown
to orangish brown at the tip. Ray florets 6–8, the corolla showy, (10–)15–30 mm
long, yellow. Disc florets 40–100(–150), the corollas 3–4 mm long, yellow to
orangish yellow. Pappus of (2–)4 awns mostly 2–4 mm long, these with downward-pointed
barbs, erect at fruiting. Fruits 4–8 mm long, narrowly wedge-shaped, more or
less flattened and somewhat 4-angled in cross-section, the angles not pale and
not thickened or winged, the faces sometimes with several faint longitudinal
lines, dark brown to purplish black, glabrous or with sparse, short,
pustular-based hairs. 2n=22, 24. August–October.
Uncommon, known
thus far only from a single historical collection from Dunklin County (southern
U.S. north to California, Missouri, and Maine; Mexico, Central America, South
America; introduced in Hawaii). Habitat unknown, but presumably banks of
streams and rivers or bottomland forests.
Steyermark
(1963) noted that a number of additional specimens originally determined as B.
laevis had been redetermined during his studies as the closely related B.
cernua. He also discussed that the two species tend to intergrade and
perhaps should be considered varieties of a single species.