5. Leucanthemum Mill.
About 33
species, Europe, Africa, introduced in the New World.
It has now
become widely accepted that the former broad circumscription of the genus Chrysanthemum
involved the recognition of an unnatural group. The most recent classifications
separate this unwieldy group into about 38 genera (Soreng and Cope, 1991;
Bremer and Humphries, 1993). Unfortunately, the characters supporting this
reclassification are mostly microscopic and/or anatomical, which has made it
difficult to write keys to the segregate genera. Chrysanthemum in the
strict sense is now confined to a few annual species native to the
Mediterranean region. It and a few other non-Missouri genera are part of a
group with florets with two different types of achenes: those of the showy,
mostly yellow ray florets strongly 3-ribbed or 3-winged; and those of the disc
florets thinner-walled and less strongly angled (sometimes 1 of the angles
winged) (Soreng and Cope, 1991). Both types of florets lack a pappus, but this
and some other defining characteristics are not individually unique to Chrysanthemum,
which can only be separated easily by a combination of different characters.
Species of Chrysanthemum in the strict sense are cultivated under names
like garland chrysanthemum and crown daisy (C. coronarium L.), corn
marigold (C. carinatum Schousb.), and tricolor chrysanthemum (C.
segetum L.).
In the other
genera, when both disc and ray florets produce fruits, these tend to be similar
in size and morphology. The familiar garden and florist’s mums are mostly
cultivars and hybrids involving a plant currently known as Dendranthema
×grandiflorum (Ramat.) Kitam. Leucanthemum belongs to a group of
about a dozen genera characterized by having resin ducts in the achene wall,
among other features. In addition to the species treated below, several other
species sometimes are grown as ornamentals. They include the Shasta daisy, L.×superbum (Bergmans ex J.W. Ingram) Soreng & E. Cope, which was
developed from a cross between the Portuguese chrysanthemum, L. lacustre
(Brot.) &Cope, 1991).