1. Amsinckia Lehm.
Plants annual,
with slender taproots. Stems erect or ascending, unbranched or branched,
pubescent with sparse to moderate, spreading, bristly, pustular-based hairs and
usually also sparse to dense, fine, short, more or less appressed, sometimes
somewhat matted hairs. Leaves alternate and often also basal, sessile,
gradually reduced toward the stem tip. Leaf blades linear to narrowly oblong or
oblong-lanceolate, the uppermost sometimes narrowly ovate, narrowed to somewhat
rounded at the base, mostly angled to a bluntly or sharply pointed tip, the
lowermost leaves tapered at the base and often rounded at the tip, the surfaces
and margins moderately to densely pubescent with more or less spreading,
bristly, pustular-based hairs. Inflorescences not paired, at first appearing as
dense, terminal and apparently axillary (terminal on short axillary branches)
clusters, subsequently elongating into ascending, scorpioid, spikelike racemes,
occasionally appearing as few-branched panicles, the flowers short-stalked,
usually lacking bracts (rarely a few present at the base of the inflorescence).
Calyces more or less actinomorphic, 5-lobed nearly to the base, occasionally
some of the lobes fused (the calyx then appearing 3- or 4-lobed), the lobes
mostly linear to narrowly lanceolate-triangular, with bristly, pustular-based
hairs, persistent and ascending at fruiting. Corollas more or less
trumpet-shaped, actinomorphic, yellow or orange, sometimes with reddish streaks
in the throat, the throat sometimes with small, scalelike appendages, the lobes
more or less spreading, rounded. Stamens attached above or less commonly below
the midpoint of the corolla tube, the filaments very short, the anthers oblong,
not exserted from the corolla. Ovary deeply 4-lobed, the style slender, not
exserted from the corolla, often not persistent at fruiting, the stigma
capitate, unlobed. Fruits usually dividing into 4 nutlets, these erect,
angular-ovoid, attached to the pyramidal gynobase toward the basal portion of a
ventral keel, the relatively small scar usually surrounded by a small outgrowth
of tissue, the surface appearing variously roughened, warty, or somewhat
wrinkled, usually at least in part with small, blunt tubercles, white to
grayish white or occasionally tan. Fifteen to 50 species, North America, South America.
The taxonomy of Amsinckia
is complex and not well understood, with species boundaries difficult to
evaluate because of confusing patterns of morphological variation between
populations. Many new species were described by Suksdorf (1931), but the
majority of them now seem unworthy of recognition. The nomenclature
consequently is confusing, with some species treated under different names in
various state floristic manuals. In the present work, the nomenclatural
suggestions of Ray and Chisaki (1957) are followed in dealing with these taxa.
In general, Amsinckia
species are weedy and are found in open, disturbed sites in a variety of soil
types. All the taxa found in Missouri
are considered toxic to livestock, especially to horses (Burrows and Tyrl,
2001). Collectors should also exercise caution as the bristly hairs found in
abundance on the plants can cause itching and/or dermatitis.