6. Senna Mill.
(senna)
Plants annual or
perennial herbs (trees and shrubs elsewhere), unarmed, with 1 to several stems,
the roots often blackish; root nodules lacking. Leaves short- to more commonly
relatively long-petiolate, the petiole or rachis base with a large, variously
shaped gland. Stipules small and scalelike to nearly hairlike, often shed
early. Leaf blades evenly 1 time pinnately compound. Leaflets few to numerous,
opposite, variously shaped, often asymmetrical at the base (with one side
angled and the other rounded or cordate), the margins entire. Inflorescences
axillary racemes, sometimes appearing aggregated in terminal clusters, the
flower stalks lacking bracts. Flowers perfect, slightly perigynous, slightly to
moderately asymmetrical, the buds usually nodding. Hypanthium very short and more
or less disc-shaped. Calyces of 5 free sepals. Corollas of 5 free petals; these
dissimilar in shape and position and usually slightly dissimilar in size,
abruptly tapered at the base, rounded at the tip, yellow to orangish yellow,
drying white, often with dark veins. Stamens 10 but the upper 3 reduced to
staminodes, the filaments short, the anthers attached at the base, graded in
size, the upper 3 strongly reduced and infertile, the middle 4 intermediate and
fertile, the lower 3 large and fertile, all somewhat curved, the fertile ones
dehiscing by an apical pore. Styles curved. Fruits legumes, elongate, not
twisted, flattened to circular or rectangular in cross-section, narrowed to a
short, stalklike base, indehiscent or, if dehiscent, then the valves not
separating elastically or coiling. Seeds variously shaped and colored, usually
somewhat flattened, with an elliptic pleurogram, this sometimes conspicuously
different in color from the remainder of the seed. About 240–260 species,
nearly worldwide, most diverse in the New World
tropics.
Senna is a large, widespread, and
morphologically diverse genus that traditionally was included in a broadly
defined Cassia L. along with Chamaecrista. The differences
between these taxa were discussed by Irwin and Barneby (1981, 1982), whose
overall classification presently is followed by most botanists. In contrast to Chamaecrista,
the stipules of Senna are small and weakly developed, and the flowers
lack small bracts on the stalks; the stamens of Senna have relatively large,
curved anthers of various sizes in a more or less graded series.