3. Triosteum perfoliatum L. (common horse gentian, wild coffee, tinkers weed,
feverwort)
Map 1436, Pl.
336 a, b
Stems 0.3–1.2 m
long, densely pubescent with short (0.3–0.5 mm), softer hairs that are mostly
minutely gland-tipped and also sparse straight, more or less spreading, longer,
bristly hairs 1.0–1.5 mm long. Median 3–5 leaf pairs usually moderately to
strongly perfoliate, these 1–6 cm wide at the base. Leaf blades 9–20 cm long, 4–9
cm wide, oblanceolate to obovate or elliptic, those of the largest leaves
occasionally slightly fiddle-shaped, the nonperfoliate leaves tapered at the
base, sometimes to an indistinct, broadly winged petiolar base, tapered to a
sharply pointed tip, the margins with relatively dense, stiff, ascending hairs,
the upper surface moderately pubescent with short, straight, appressed hairs,
the undersurface moderately to densely pubescent with short, soft, spreading
hairs mostly along the veins. Flowers 1 or more commonly 2 or 3(4) per leaf
axil (2–6[8] per node). Paired bracts subtending each flower sometimes longer
than the ovary but shorter than the flower (including the calyx and corolla),
linear to narrowly lanceolate. Calyx lobes 10–18 mm long, the margins with
dense, stiff, bristly, longer and shorter hairs, the inner and outer surfaces
with scattered, short hairs, some of these often minutely gland-tipped.
Corollas 8–17 mm long, dull red to purplish red, often mottled with small,
cream-colored spots near the base and on the inner surface, occasionally
entirely greenish yellow, narrowly funnelform, the mouth only slightly oblique,
the outer surface with gland-tipped hairs. Styles usually slightly to
moderately exserted (to 5 mm beyond the corolla lobes). Fruits 7–10 mm in
diameter, orangish yellow to orange, sparsely hairy at maturity. May–July.
Scattered nearly
throughout the state, most abundantly in the Ozark and Ozark Border Divisions
(eastern U.S. west to Minnesota, Nebraska, and Texas; Canada). Bottomland
forests, mesic upland forests, bases and ledges of bluffs, edges of glades, and
banks of streams and rivers; also roadsides.
This is the most
abundant and widespread member of the genus in Missouri. See the treatment of T.
aurantiacum for a discussion of problems in discriminating that species
from T. perfoliatum.