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  • Reference article Neupane, S., S. Dessein, N. Wikström, P. O. Lewis, C. Long, B. Bremer & T. J. Motley. 2015. The Hedyotis-Oldenlandia complex (Rubiaceae, Spermaacoceae) in Asia and the Pacific: Phylogeny revisited with new generic delimitations. Taxon 64(2): 299–322.
 
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