Annual or perennial caespitose herbs. Leaves altetnate. Flowers in simple or branched terminal racemes. Calyx 5-partite, sometimes accrescent and reflexed in fruit. Corolla funnel-form to ± rotate; lobes spreading; throat appendages present. Stamens included. Style filiform, stigma capitate. Nutlets 4, with margins appendiculate, smooth to rugose or tuberculate.