Involucre phyllaries usually with prominent subterminal horns (corniculate to weakly bifurcate or cornute to prominently bifurcate), rarely smooth or callose. Cypselas 3 – 5 mm long, greenish, brown, stramineous, orangish or reddish-brown with usually 1 or more than 1 mm long, cylindrical cone formed abruptly, rarely small in T. pullocarpum; beak long; pappus white to usually snow white.