Frangula capreifolia var. grandifolia (M. C. Johnst. & L. A. Johnst.) A. Pool, ined.; Rhamnus capreifolia var. grandifolia M. C. Johnst. & L. A. Johnst.
Usually small trees (possibly shrub), (3--) 6—13 (--23) m, young branches sparse to dense pilose, with golden trichomes, or less frequently glabrous. Leaves with blades 7--20 x (2.8--) 3.5—8.5 cm, elliptic, or less frequently, oblanceolate, membranaceous, both surfaces of similar color, 8--14 pairs of lateral veins, lateral veins at 45–75 degree angle to midrib, adaxial surface with midrib and lateral veins pilose or with appressed to ascending trichomes, surface otherwise glabrous or with scattered, appressed to ascending trichomes, abaxial surface with numerous (rarely few) trichomes more or less straight and extending laterally from sides of midrib and often lateral veins, often macroscopically conspicuously concentrated along sides of midrib and/or in angles of lateral veins, venation also sometimes pilose, the trichomes sometimes in tufts of 2—4 from one point, base cuneate to rounded, sometime inequilateral, margin flat to recurved with 3—6 (--8) teeth/cm, the teeth overlapping at base over adaxial surface or often only apical gland apparent, apex acuminate or acuminate-falcate (rarely acute or obtuse), tip often mucronate; stipules very early caducous, 2.75—4.5 mm; petiole 10--25 mm. Inflorescence a sessile fascicle, a pedunculate and umbelliform cyme, or a pedunculate compound cyme with one arm of cyme umbelliform and the other arm continuing as a rachis which develops another umbelliform cluster, each umbelliform cluster or fascicle with 6--20 flowers; peduncle 0—15 (--25) mm, 1/5 to ca. equal in length to subtending petiole (similar in fruit; rarely to 2 times as long). Flowers on pedicels 2.5—10 mm, 1/5 to 5 times length of peduncle when present, 1/5--2/3rds (--5/6ths) length of subtending petiole (similar in fruit); flower 2—3.5 mm, hypanthium campanulate, 1.4—2 x 2—3.5 (4) mm, hypanthium width (rarely 2/3) ca. equal to 1.5 times flower length, sparse to abundant pilose, with spreading trichomes, circumscissile. Sepals, petals and stamens lost early with upper portion of hypanthium. Sepals 0.75—1.5 (--2) mm long and wide, slightly shorter than to less frequently equal to hypanthium in length, spreading or recurved. Petals 0.75—1.3 (--1.6) mm long and wide, weakly to strongly apically bi-lobed, glabrous or midrib abaxially puberulent, slightly exserted; stamens with filaments 0.7—1 mm, anthers 0.5—0.75 mm, infrequently visible free from petals. Ovary with dense long trichomes, style 0.75—1.75 mm, stigma 3-lobed. Young fruits 1—8 (13) fascicled, with few to numerous trichomes, globose or obovoid, 3 lines conspicuous; all sepals, petals and stamen filaments rapidly lost, hypanthium remnant 0.5—1 x 2—3 mm, not angled; stones 3, ca. 6 x 5 mm.
Expected in Nicaragua at higher elevations, known from Mexico, Guatemla, El Salvador, and Costa Rica. Treated in the published
Flora de Nicaragua as
Rhamnus capraefolia var.
grandifolia. The illustration labeled Rhamnus “capreaefolia” in the Flora of Panama depicts Frangula discolor var. discolor.